Itwasthereforeinnearlythecertaintyofnon-fulfilmentthatImademyfirstattemptsatrearingcarnivorouslarvaewithaquarrywhichdidnotconformwiththecustomaryregimen。Forconscience\'sake,moreorlessperfunctorily,Iendeavouredtoachievesomethingthatseemedtomeboundtoendinpitifulfailure。OnlytheBembex-wasps,whichareplentifulinthesandoftheneighbouringhills,mightstillaffordme,withouttooprolongedasearch,afewsubjectsonwhichtoexperiment。TheTarsalBembexfurnishedmewithwhatIwanted:larvaeyoungenoughtohavestillbeforethemalongperiodoffeedingandyetsufficientlydevelopedtoendurethetrialsofaremoval。
Theselarvaareexhumedwithalltheconsiderationwhichtheirdelicateskindemands;anumberofheadofgamearelikewiseunearthedintact,havingbeenrecentlybroughtbythemother。TheyconsistofvariousDiptera,includingsomeAnthrax-flies。(Cf。“TheLifeoftheFly“:chapters2and4——Translator\'sNote。)Anoldsardine-box,containingalayerofsiftedsandanddividedintocompartmentsbypaperpartitions,receivesmycharges,whoareisolatedonefromanother。TheseFly-eatersIproposetoturnintoGrasshopper-eaters;fortheirBembex-dietIintendtosubstitutethedietofaSphexoraTachytes。Tosavemyselftediouserrandsdevotedtoprovisioningtherefectory,Iacceptwhatgoodfortuneoffersmeattheverythresholdofmydoor。AgreenLocustid,withashortsabrebentintoareaping-hook,Phaneropterafalcata,isravagingthecorollaeofmypetunias。Nowisthetimetoindemnifymyselfforthedamagewhichshehascausedme。Ipickheryoung,halftothree-quartersofaninchinlength;
andIdepriveherofmovement,withoutmoreado,bycrushingherhead。InthisconditionsheisserveduptotheBembex-larvaeinplaceoftheirFlies。
Ifthereaderhassharedmyconvictionsoffailure,convictionsbasedonverylogicalmotives,hewillnowsharemyprofoundsurprise。Theimpossiblebecomespossible,thesenselessbecomesreasonableandtheexpectedbecomestheoppositeofthereal。ThedishservedontheBembeces\'
tableforthefirsttimesinceBembecescameintotheworldisacceptedwithoutanyrepugnanceandconsumedwitheverymarkofsatisfaction。Iwillheresetdownthedetaileddiaryofoneofmyguests;thatoftheotherswouldonlybearepetition,saveforafewvariations。
2AUGUST,1883——ThelarvaoftheBembex,asIextractitfromitsburrow,isabouthalf-developed。ArounditIfindonlysomescantyrelicsofitsmeals,consistingchieflyofAnthrax-wings,half-diaphanousandhalf-
clouded。Themotherwouldappeartohavecompletedthevictuallingbyfreshcontributions,addeddaybyday。Igivethenurseling,whichisanAnthrax-
eater,ayoungPhaneroptera。TheLocustidisattackedwithouthesitation。
Thisprofoundchangeinthecharacterofitsvictualsdoesnotseemintheleasttodisturbthelarva,whichbitesstraightintotherichmorselwithitsmandiblesanddoesnotletgountilithasexhaustedit。Towardseveningthedrainedcarcaseisreplacedbyanother,quitefresh,ofthesamespeciesbutbulkier,measuringoverthree-quartersofaninch。
3AUGUST——NextdayIfindthePhaneropteradevoured。Nothingremainsbutthedryinteguments,whicharenotdismembered。Theentirecontentshavedisappeared;thegamehasbeenemptiedthroughalargeopeningmadeinthebelly。AregularGrasshopper-eatercouldnothaveoperatedmoreskilfully。
IreplacetheworthlesscarcasebytwosmallLocustidae。Atfirstthelarvadoesnottouchthem,beingamplysatedwiththecopiousmealofthedaybefore。Intheafternoon,however,oneoftheitemsisresolutelyattacked。
4AUGUST——Irenewthevictuals,althoughthoseofthedaybeforearenotfinished。Fortherest,Idothesamedaily,sothatmychargemayconstantlyhavefreshfoodathand。Highgamemightupsetitsstomach。MyLocustidaearenotvictimsatthesametimelivingandinert,operateduponaccordingtothedelicatemethodoftheinsectsthatparalysetheirprey;
theyarecorpses,procuredbyabrutalcrushingofthehead。Withthetemperaturenowprevailing,fleshsoonbecomestainted;andthiscompelsmefrequentlytorenewtheprovisionsinmysardine-boxrefectory。Twospecimensareservedup。Oneisattackedsoonafterwards;andthelarvaclingstoitassiduously。
5AUGUST——Theravenousappetiteofthestartisbecomingassuaged。Mysuppliesmaywellbetoogenerous;anditmightbeprudenttotryalittledietingafterthisGargantuangoodcheer。Themothercertainlyismoreparsimonious。Ifallthefamilyweretoeatatthesamerateasmyguest,shewouldneverbeabletokeeppacewiththeirdemands。Therefore,forreasonsofhealth,thisisadayoffastingandvigil。
6AUGUST——SuppliesarerenewedwithtwoPhaneropterae。Oneisconsumedentirely;theotherisbitteninto。
7August——To-day\'srationistastedandthenabandoned。Thelarvaseemsuneasy。Withitspointedmouthitexploresthewallsofitschamber。Thissigndenotestheapproachofthetimeformakingthecocoon。
8AUGUST——Duringthenightthelarvahasspunitssilkeneel-trap。Itisnowencrustingitwithgrainsofsand。Thenfollow,induetime,thenormalphasesofthemetamorphosis。FedonLocustidae,adietunknowntoitsrace,thelarvapassesthroughitsseveralstageswithoutanymoredifficultythanitsbrothersandsistersfedonFlies。
IobtainedthesamesuccessinofferingyoungMantesforfood。Oneofthelarvaethusservedwouldeveninclinemetobelievethatitpreferredthenewdishtothetraditionaldietofitsrace。TwoEristales,orDrone-
flies,andaPrayingMantisaninchlongcomposeditsdailyallowance。TheDrone-fliesaredisdainedfromthefirstmouthful;andtheMantis,alreadytastedandapparentlyfoundexcellent,causestheFlytobecompletelyforgotten。Isthisanepicure\'spreference,duetothegreaterjuicinessoftheflesh?Iamnotinapositiontosay。Atallevents,theBembexisnotsoinfatuatedwithFlyastorefusetoabandonitforothergame。
ThefailurewhichIforesawhasprovedamagnificentsuccess。Itisfairlyconvincing,isitnot?Withouttheevidenceofexperiment,whatcanwerelyupon?BeneaththeruinsofsomanytheorieswhichappearedtobemostsolidlyerectedIshouldhesitatetoadmitthattwoandtwomakefourifthefactswerenotbeforeme。Myargumenthadthemosttemptingprobabilityonitsside,butithadnotthetruth。Asitisalwayspossibletofindreasonsaftertheeventinsupportofanopinionwhichonewouldnotatfirstadmit,Ishouldnowargueasfollows:
Theplantisthegreatfactoryinwhichareelaborated,withmineralmaterials,theorganicprincipleswhicharethematerialsoflife。Certainproductsarecommontothewholevegetableseries,butothers,farlessnumerous,arepreparedinspeciallaboratories。Eachgenus,eachspecieshasitstrade-mark。Hereessentialoilsaremanufactured;herealkaloids;
herestarches,fattysubstances,resins,sugars,acids。Henceresultspecialenergies,whichdonotsuiteveryherbivorousanimal。Itassuredlyrequiresastomachmadeexpresslyforthepurposetodigestaconite,colchicum,hemlockorhenbane;thosewhohavenotsuchastomachcouldneverendureadietofthatsort。Besides,theMithridatesfedonpoisonresistonlyasingletoxin。(MithridatesVI。KingofPontus(d。B。C。63)issaidtohavesecuredimmunityfrompoisonbytakingincreaseddosesofit——
Translator\'sNote。)ThecaterpillaroftheDeath\'s-headHawk-moth,whichdelightsinthesolaninofthepotato,wouldbekilledbytheacridprincipleofthetithymalsthatformthefoodoftheSpurge-caterpillar。
Theherbivorouslarvaearethereforeperforceexclusiveintheirtastes,becausedifferentgeneraofvegetablespossessverydifferentproperties。
Withthisvarietyintheproductsoftheplant,theanimal,aconsumerfarmorethanaproducer,contraststheuniformityinitsownproducts。ThealbumenintheeggoftheOstrichortheChaffinch,thecaseininthemilkoftheCowortheAss,themuscularfleshoftheWolfortheSheep,theScreech-owlortheField-mouse,theFrogortheEarth-worm:theseremainalbumen,caseinorfibrin,edibleifnoteaten。Herearenoexcruciatingcondiments,nospecialacridities,noalkaloidsfataltoanystomachotherthanthatoftheappointedconsumer;sothatanimalfoodisnotconfinedtooneandthesameeater。Whatdoesnotmaneat,fromthatdelicacyofthearcticregions,soupmadeofSeal\'sbloodandascrapofWhale-blubberwrappedinawillow-leafforavegetable,totheChinaman\'sfriedSilk-wormortheArab\'sdriedLocust?Whatwouldhenoteat,ifhehadnottoovercometherepugnancedictatedbyhabitratherthanbyactualnecessity?
Thepreybeinguniforminitsnutritiveprinciples,thecarnivorouslarvaoughttoaccommodateitselftoanysortofgame,aboveallifthenewdishbenottoogreatadeparturefromconsecratedusage。ThusshouldIargue,withnolessprobabilityonmyside,hadItobeginalloveragain。But,asallourargumentshavenotthevalueofasinglefact,Ishouldbeforcedintheendtoresorttoexperiment。
Ididsothenextyear,onalargerscaleandwithagreatervarietyofsubjects。Ishrinkfromacontinuousnarrativeofmyexperimentsandofmypersonaleducationinthisnewart,wherethefailureofonedaytaughtmethewaytosucceedonthemorrow。Itwouldbelongandtedious。EnoughifI
brieflystatemyresultsandtheconditionswhichmustbefulfilledinordertorunthedelicaterefectoryasitshouldberun。
And,first,wemustnotdreamofdetachingtheeggfromitsnaturalpreytolayitonanother。Theeggadheresprettyfirmly,byitscephalicpole,tothequarry。Toremoveitfromitsplacewouldinevitablyjeopardizeitsfuture。Ithereforeletthelarvahatchandacquiresufficientstrengthtobeartheremovalwithoutperil。Forthatmatter,myexcavationsmostoftenprovidemewithmysubjectsintheformoflarvae。Iadoptforrearing-
purposesthelarvaethatareaquartertoahalfdeveloped。Theothersaretooyoungandriskytohandle,ortoooldandlimitedtoashortperiodofartificialfeeding。
Secondly,Iavoidbulkyheadsofgame,asingleoneofwhichwouldsufficeforthewholegrowing-stage。IhavealreadysaidandIhererepeathowniceamatteritistoconsumeavictimwhichhastokeepfreshforacoupleofweeksandnottofinishdyinguntilitisalmostentirelydevoured。Deathhereleavesnocorpse;whenlifeisextinct,thebodyhasdisappeared,leavingonlyashredofskin。Larvaewithonlyonelargepreyhaveaspecialartofeating,adangerousart,inwhichaclumsybitewouldprovefatal。Ifbittenbeforethepropertimeatsuchapoint,thevictimbecomesputrid,whichpromptlycausesdeathbypoisoningintheconsumer。Whendivertedfromitsplanofattack,deprivedofitsclue,thelarvaisnotalwaysabletorediscoverthelawfulmorselsingoodtimeandiskilledbythedecompositionofitsbadlydissectedprey。Whatwillhappeniftheexperimentergivesitagametowhichitisnotaccustomed?Notknowinghowtoeatitaccordingtorule,thelarvawillkillit;andbynextdaythevictualswillhavebecomesomuchtoxicputrescence。IhavealreadytoldhowIfounditimpossibletoreartheTwo-bandedScoliaonOryctes-larvae,fasteneddowntodeprivethemofmovement,orevenonEphippigers,paralysedbytheLanguedocianSphex。Inbothcasesthenewdietwasacceptedwithouthesitation,aproofthatitsuitedthenurseling;butinadayortwoputrescencesupervenedandtheScoliaperishedonthefetidmorsel。ThemethodofpreservingtheEphippiger,sowellknowntotheSphex,wasunknowntomyboarder;inthiswasenoughtoconvertadeliciousfoodintopoison。
Evensodidmyotherattemptsmiscarrywretchedly,attemptsatfeedingwiththesingledishconsistingofonebigheadofgametoreplacethenormalration。Onlyonesuccessisrecordedinmynotebooks,butthatwassodifficultthatIwouldnotundertaketoobtainitasecondtime。I
succeededinfeedingthelarvaoftheHairyAmmophilawithanadultblackCricket,whowasacceptedasreadilyasthenaturalgame,thecaterpillar。
Toavoidputrefactionofvictualswhichlastoverlongandarenotconsumedaccordingtothemethodindispensabletotheirpreservation,Iemploysmallgame,eachpieceofwhichcanbefinishedbythelarvaatasinglesitting,oratmostinasingleday。Itmatterslittlethenthatthevictimisslashedanddismemberedatrandom;decompositionhasnotimetoseizeuponitsstillquiveringtissues。Thisistheprocedureofthoselarvaewhichgulpdowntheirfood,snappingatrandomwithoutdistinguishingonepartfromanother,suchastheBembex-larvae,whichfinishtheFlyintowhichtheyhavebittenbeforebeginninganotherintheheap,ortheCerceris-
larvae,whichdraintheirWeevilsmethodicallyoneafteranother。Withthefirststrokesofthemandiblesthevictimbroachedmaybemortallywounded。
Thisisnodisadvantage:abriefspellsufficestomakeuseofthecorpse,whichissavedfromputrefactionbybeingpromptlyconsumed。Closebesideit,theothervictims,quitealivethoughmotionless,awaittheirrespectiveturnsandsupplyreservesofvictualswhicharealwaysfresh。
IamtoounskilfulabutchertoimitatetheWaspandmyselftoresorttoparalysis;moreover,thecausticliquidinjectedintothenerve-centres,ammoniainparticular,wouldleavetracesofsmellorflavourwhichmightputoffmyboarders。Iamthereforecompelledtodeprivemyinsectsofthepowerofmovementbykillingthemoutright。Thismakesitimpracticabletoprovideasufficiencyofprovisionsbeforehandinasinglesupply:whileoneitemoftherationwasbeingconsumedtherestwouldspoil。Oneexpedientaloneremainstome,onewhichentailsconstantattendance:itistorenewtheprovisionseachday。Whenalltheseconditionsarefulfilled,thesuccessofartificialfeedingisstillnotwithoutitsdifficulties;
nevertheless,withalittlecareandaboveallplentyofpatience,itisalmostcertain。
ItwasthusthatIrearedtheTarsalBembex,whicheatsAnthrax-fliesandotherDiptera,onyoungLocustidaeorMantidae;theSilkyAmmophila,whosedietconsistschieflyofMeasuring-worms,onsmallSpiders;thepot-makingPelopaeus,aSpider-eater,ontenderAcridians;theSandCerceris,apassionateloverofWeevils,onHalicti;theBee-eatingPhilanthus,whichfeedsexclusivelyonHive-bees,onEristalesandotherFlies。Withoutsucceedinginmyfinalaim,forreasonswhichIhavejustexplained,IhaveseentheTwo-bandedScoliafeastinggreedilyonthegruboftheOryctes,whichwassubstitutedforthatoftheCetonia,andputtingupwithanEphippigertakenfromtheburrowoftheSphex;IhavebeenpresentattherepastofthreeHairyAmmophilaeacceptingwithanexcellentappetitetheCricketthatreplacedtheircaterpillar。Oneofthem,asIhaverelated,contrivedtokeepitsrationfresh,whichenabledittoreachitsfulldevelopmentandtospinitscocoon。
Theseexamples,theonlyonestowhichmyexperimentshaveextendedhitherto,seemtomesufficientlyconvincingtoallowmetoconcludethatthecarnivorouslarvadoesnothaveexclusivetastes。Therationsuppliedtoitbythemother,somonotonous,solimitedinquality,mightbereplacedbyothersequallytoitstaste。Varietydoesnotdispleasethelarva;itdoesitasmuchgoodasuniformity;indeed,itwouldbeofgreaterbenefittotherace,asweshallseepresently。
CHAPTER8。ADIGATTHEEVOLUTIONISTS。
Torearacaterpillar-eateronaskewerfulofSpidersisaveryinnocentthing,unlikelytocompromisethesecurityoftheState;itisalsoaverychildishthing,asIhastentoconfess,andworthyoftheschoolboywho,inthemysteriesofhisdesk,seeksasbesthemaysomediversionfromthefascinationsofhisexerciseincomposition。AndIshouldnothaveundertakentheseinvestigations,stilllessshouldIhavespokenthem,notwithoutsomesatisfaction,ifIhadnotdiscerned,intheresultsobtainedinmyrefectory,acertainphilosophicimport,involving,soitseemedtome,theevolutionarytheory。
Itisassuredlyamajesticenterprise,commensuratewithman\'simmenseambitions,toseektopourtheuniverseintothemouldofaformulaandsubmiteveryrealitytothestandardofreason。Thegeometricianproceedsinthismanner:hedefinesthecone,anidealconception;thenheintersectsitbyaplane。Theconicsectionissubmittedtoalgebra,anobstetricalappliancewhichbringsforththeequation;andbehold,entreatednowinonedirection,nowinanother,thewomboftheformulagivesbirthtotheellipse,thehyperbola,theparabola,theirfoci,theirradiusvectors,theirtangents,theirnormals,theirconjugateaxes,theirasymptotesandtherest。Itismagnificent,somuchsothatyouareovercomebyenthusiasm,evenwhenyouaretwentyyearsold,anagehardlyadaptedtotheausteritiesofmathematics。Itissuperb。Youfeelasifyouwerewitnessingthecreationofaworld。
Asamatteroffact,youaremerelyobservingthesameideafromdifferentpointsofview,whichareilluminedbythesuccessivephasesofthetransformedformula。Allthatalgebraunfoldsforourbenefitwascontainedinthedefinitionofthecone,butitwascontainedasagerm,underlatentformswhichthemagicofthecalculusconvertsintoexplicitforms。Thegrossvaluewhichourmindconfidedtotheequationitreturnstous,withoutlossorgain,incoinsstampedwitheverysortofeffigy。Andherepreciselyisthatwhichconstitutestheinflexiblerigourofthecalculus,theluminouscertaintybeforewhicheverycultivatedmindisforcedtobow。
Algebraistheoracleoftheabsolutetruth,becauseitrevealsnothingbutwhatthemindhadhiddeninitunderanamalgamofsymbols。Weput2and2
intothemachine;therollersworkandshowus4。Thatisall。
Buttothiscalculus,all-powerfulsolongasitdoesnotleavethedomainoftheideal,letussubmitaverymodestreality:thefallofagrainofsand,thependularmovementofahangingbody。Themachinenolongerworks,ordoessoonlybysuppressingalmosteverythingthatisreal。Itmusthaveanidealmaterialpoint,anidealrigidthread,anidealpointofsuspension;andthenthependularmovementistranslatedbyaformula。Buttheproblemdefiesalltheartificesofanalysisiftheoscillatingbodyisarealbody,endowedwithvolumeandfriction;ifthesuspensorythreadisarealthread,endowedwithweightandflexibility;ifthepointofsupportisarealpoint,endowedwithresistanceandcapableofdeflection。Sowithotherproblems,howeversimple。Theexactrealityescapestheformula。
Yes,itwouldbeafinethingtoputtheworldintoanequation,toassumeasthefirstprincipleacellfilledwithalbumenandbytransformationaftertransformationtodiscoverlifeunderitsthousandaspectsasthegeometriciandiscoverstheellipseandtheothercurvesbyexamininghisconicsection。Yes,itwouldbemagnificentandenoughtoaddacubittoourstature。Alas,howgreatlymustweabateourpretensions!Therealityisbeyondourreachwhenitisonlyamatteroffollowingagrainofdustinitsfall;andwewouldundertaketoascendtheriveroflifeandtraceittoitssource!Theproblemisamorearduousonethanthatwhichalgebradeclinestosolve。Thereareformidableunknownquantitieshere,moredifficulttodecipherthantheresistances,thedeflectionsandthefrictionsofthependulum。Letuseliminatethem,thatwemaymoreeasilypropoundthetheory。
Verywell;butthenmyconfidenceinthisnaturalhistorywhichrepudiatesnatureandgivesidealconceptionsprecedenceoverrealfactsisshaken。
So,withoutseekingtheopportunity,whichisnotmybusiness,Itakeitwhenitpresentsitself;Iexaminethetheoryofevolutionfromeveryside;
and,asthatwhichIhavebeenassuredisthemajesticdomeofamonumentcapableofdefyingtheagesappearstometobenomorethanabladder,I
irreverentlydigmypinintoit。
Hereisthelatestdig。Adaptabilitytoavarieddietisanelementofwell-beingintheanimal,afactorofprimeimportancefortheextensionandpredominanceofitsraceinthebitterstruggleforlife。Themostunfortunatespecieswouldbethatwhichdependedforitsexistenceonadietsoexclusivethatnoothercouldreplaceit。WhatwouldbecomeoftheSwallowifherequired,inordertolive,oneparticularGnat,asingleGnat,alwaysthesame?WhenoncethisGnathaddisappeared——andthelifeoftheMosquitoisnotalongone——thebirdwoulddieofstarvation。
Fortunatelyforhimselfandforthehappinessofourhomes,theSwallowgulpsthemalldownindiscriminately,togetherwithahostofotherinsectsthatperformaerialballets。WhatwouldbecomeoftheLarkwerehisgizzardabletodigestonlyoneseed,invariablythesame?Whentheseasonforthisseedwasover——andtheseasonisalwaysashortone——thehaunterofthefurrowswouldperish。
Isnotman\'scomplaisantstomach,adaptedtothelargestvarietyofnourishment,oneofhisgreatzoologicalprivileges?Heisthusrenderedindependentofclimates,seasonsandlatitudes。AndtheDog:howisitthatofallthedomesticanimalshealoneisabletoaccompanyuseverywhere,evenonthemostarduousexpeditions?TheDogagainisomnivorousandthereforeacosmopolitan。
Thediscoveryofanewdish,saidBrillat-Savarin,isofgreaterimportancetohumanitythanthediscoveryofanewplanet。Theaphorismisnearertothetruththanitappearstobeinitshumorousform。Certainlythemanwhowasthefirsttothinkofcrushingwheat,kneadingflourandcookingthepastebetweentwohotstoneswasmoredeservingthanthediscovererofthetwo-hundredthasteroid。TheinventionofthepotatoiscertainlyasvaluableasthatofNeptune,gloriousasthelatterwas。Allthatincreasesouralimentaryresourcesisadiscoveryofthefirstmerit。Andwhatistrueofmancannotbeotherthantrueofanimals。Theworldbelongstothestomachwhichisindependentofspecialities。Thistruthisofthekindthathasonlytobestatedtobeproved。
Letusnowreturntoourinsects。IfIamtobelievetheevolutionists,thevariousgame-huntingWaspsaredescendedfromasmallnumberoftypes,whicharethemselvesderived,byanincalculablenumberofconcatenations,fromafewamoebae,afewmoneraandlastlyfromthefirstclotofprotoplasmwhichwascasuallycondensed。Letusnotgobackasfarasthat;
letusnotplungeintothefogswhereillusionanderrortooeasilyfindalurking-place。Letusconsiderasubjectwithexactlimitstoit;thisistheonlywaytounderstandoneanother。
TheSphegidaearedescendedfromasingletype,whichitselfwasalreadyahighly-developeddescendantand,likeitssuccessors,feditsfamilyonprey。Theclosesimilarityinform,incolouringand,aboveall,inhabitsseemtorefertheTachytestothesameorigin。Thisisample;letusbesatisfiedwithit。Andnowpleasetellme,whatdidthisprototypeoftheSphegidaehunt?Wasitsdietvariedoruniform?Ifwecannotdecide,letusexaminethetwocases。
Thedietwasvaried。IheartilycongratulatethefirstbornoftheSphex-
wasps。Sheenjoyedthemostfavourableconditionsforleavingaprosperousoffspring。Accommodatingherselftoanykindofpreynotdisproportionatetoherstrength,sheavoidedthedearthofagivenspeciesofgameatthisorthattimeandinthisorthatplace;shealwaysfoundthewherewithaltoendowherfamilymagnificently,theybeing,forthatmatter,fairlyindifferenttothenatureofthevictuals,providedthattheseconsistedoffreshinsect-flesh,asthetastesoftheircousinsmanytimesremovedprovetothisday。ThismatriarchoftheSphexclanborewithinherselfthebestchancesofassuringvictorytoheroffspringinthatpitilessfightforexistencewhicheliminatestheweaklyandincapableandallowsnonebutthestrongandindustrioustosurvive;shepossessedanaptitudeofgreatvaluewhichatavismcouldnotfailtohanddownandwhichherdescendants,whoaregreatlyinterestedinpreservingthismagnificentinheritance,musthavepermanentlyadoptedandevenaccentuatedfromonegenerationtothenext,fromonebranch,oneoffshoot,toanother。
Insteadofthisunscrupulouslyomnivorousrace,levyingbootyuponeverykindofgame,toitsverygreatadvantage,whatdoweseeto-day?EachSphexisstupidlylimitedtoanunvaryingdiet;shehuntsonlyonekindofprey,thoughherlarvaacceptsthemall。OnewillhavenothingbuttheEphippigerandmusthaveafemaleatthat;anotherwillhavenothingbuttheCricket。ThisonehuntstheLocustandnothingelse;thatonetheMantisandtheEmpusa。YetanotherisaddictedtotheGreyWormandanothertotheLooper。
Fools!Howgreatwasyourmistakeinallowingthewiseeclecticismofyourancestress,whoserelicsnowreposeinthehardmudofsomelacustrianstratum,tobecomeobsolete!Howmuchbetterwouldthingsbeforyouandyours!Abundanceisassured;painfulandoftenfruitlesssearchesareavoided;thelarderiscrammedwithoutbeingsubjecttotheaccidentsoftime,placeandclimate。WhenEphippigersrunshort,youfallbackuponCrickets;whentherearenoCrickets,youcaptureGrasshoppers。Butno,mybeautifulSphex-wasps,youwerenotsuchfoolsasthat。Ifinourdaysyouareeachconfinedtoastandingfamily-dish,itisbecauseyourancestressofthelacustrianschistsnevertaughtyouvariety。
Couldshehavetaughtyouuniformity?LetussupposethattheSphexofantiquity,anoviceinthegastronomicart,preparedherpottedmeatswithasinglekindofgame,nomatterwhat。Itwasthenherdescendantswho,subdividedintogroupsandconstitutedintosomanydistinctspeciesbytheslowtravailofthecenturies,realizedthatinadditiontotheancestralfarethereexistedahostofotherfoods。Traditionbeingabandoned,therewasnothingtoguidetheirchoice。Theythereforetriedabitofeverythinginthewayofinsectgame,athap-hazard;andeachtimethelarva,whosetastesalonehadtobeconsulted,wassatisfiedwiththefoodsupplied,asitisto-dayintherefectoryprovisionedbymycare。
Everyattemptledtotheinventionofanewdish,animportantevent,accordingtothemasters,aninestimableresourceforthefamily,whoweretherebydeliveredfromthemenaceofdeathandenabledtothriveoverlargeareaswhencetheabsenceorrarityofauniformgamewouldhaveexcludedit。And,aftermakinguseofahostofdifferentviandsinordertoattaintheculinaryvarietywhichisto-dayadoptedbythewholeoftheSphexnation,loandbehold,eachspeciesconfinesitselftoasinglesortofgame,outsidewhicheveryspecimenisobstinatelyrefused,notattable,ofcourse,butinthehunting-field!Byyourexperiments,fromagetoage,tohavediscoveredvarietyindiet;tohavepractisedit,tothegreatadvantageofyourrace,andtoendupwithuniformity,thecauseofdecadence;tohaveknowntheexcellentandtorepudiateitforthemiddling:oh,mySphex-wasps,itwouldbestupidifthetheoryofevolutionwerecorrect!
Toavoidinsultingyouandalsofromrespectforcommonsense,Ipreferthereforetobelievethat,ifinourdaysyouconfineyourhuntingtoasinglekindofgame,itisbecauseyouhaveneverknownanyother。Iprefertobelievethatyourcommonancestress,yourprecursor,whetherhertastesweresimpleorcomplex,isapurechimera,for,iftheywereanyrelationshipbetweenyou,havingtestedeverythinginordertoarriveattheactualfoodofeachspecies,havingeateneverythingandfounditgratefultothestomach,youwouldnow,fromfirsttolast,beunprejudicedconsumers,omnivorousprogressives。Iprefertobelieve,inshort,thatthetheoryofevolutionispowerlesstoexplainyourdiet。Thisistheconclusiondrawnfromthedining-roominstalledinmyoldsardine-box。
CHAPTER9。RATIONINGACCORDINGTOSEX。
Consideredinrespectofquality,thefoodhasjustdisclosedourprofoundignoranceoftheoriginsofinstinct。Successfallstotheblusterers,totheimperturbabledogmatists,fromwhomanythingisacceptedifonlytheymakealittlenoise。Letusdiscardthisbadhabitandadmitthatreally,ifwegotothebottomofthings,weknownothingaboutanything。
Scientificallyspeaking,natureisariddletowhichhumancuriosityfindsnodefinitesolution。Hypothesisfollowshypothesis;thetheoreticalrubbish-heapgrowsbiggerandbigger;andstilltruthescapesus。Toknowhowtoknownothingmightwellbethelastwordofwisdom。
Consideredinrespectofquantity,thefoodsetsusotherproblems,nolessobscure。Thoseofuswhodevoteourselvesassiduouslytostudyingthecustomsofthegame-huntingWaspssoonfindourattentionarrestedbyaveryremarkablefact,atthetimewhenourmind,refusingtobesatisfiedwithsweepinggeneralities,whichourindolencetooreadilymakesshiftwith,seekstoenterasfaraspossibleintothesecretofthedetails,socuriousandsometimessoimportant,asandwhentheybecomebetter-knowntous。Thisfact,whichhaspreoccupiedmeformanyalongyear,isthevariablequantityoftheprovisionspackedintotheburrowasfoodforthelarva。
Eachspeciesisscrupulouslyfaithfultothedietofitsancestors。FormorethanaquarterofacenturyIhavebeenexploringmydistrict;andI
haveneverknownthediettovary。To-day,asthirtyyearsago,eachhuntressmusthavethegamewhichIfirstsawherpursuing。But,thoughthenatureofthevictualsisconstant,thequantityisnotso。Inthisrespectthedifferenceissogreatthathewouldneedtobeaverysuperficialobserverwhoshouldfailtoperceiveitonhisfirstexaminationoftheburrows。Inthebeginning,thisdifference,involvingtwo,three,fourtimesthequantityandmore,perplexedmeextremelyandledmetotheconclusionswhichIrejectto-day。
Here,amongtheinstancesmostfamiliartome,aresomeexamplesofthesevariationsinthenumberofvictimsprovidedforthelarva,victims,ofcourse,verynearlyidenticalinsize。InthelarderoftheYellow-wingedSphex,afterthevictuallingiscompletedandthehouseshutup,twoorthreeCricketsaresometimesfoundandsometimesfour。Stizusruficornis(Cf。“TheHuntingWasps“:chapter20;also“Bramble-beesandOthers“:
chapter9——Translator\'sNote。),establishedinsomeveinofsoftsandstone,placesthreePrayingMantesinonecellandfiveinanother。OfthecasketsfashionedbyAmedeus\'Eumenes(Cf。“TheMason-wasps“:chapter1——Translator\'sNote。)outofclayandbitsofstone,themorerichlyendowedcontaintensmallcaterpillars,themorepoorlyfurnishedfive。TheSandCerceris(Cf。“TheHuntingWasps“:chapter2——Translator\'sNote。)
willsometimesprovidearationofeightWeevilsandsometimesoneoftwelveorevenmore。Mynotesaboundinabstractsofthiskind。Itisunnecessaryforthepurposeinhandtoquotethemall。ItwillserveourobjectbetterifIgivethedetailedinventoryoftheBee-eatingPhilanthusandoftheMantis-huntingTachytes,consideredespeciallywithregardtothequantityofthevictuals。
TheslayerofHive-beesisfrequentlyinmyneighbourhood;andIcanobtainfromherwiththeleasttroublethegreatestnumberofdata。InSeptemberI
seetheboldfilibusterflyingfromclumptoclumpofthepinkheatherpillagedbytheBee。Thebanditsuddenlyarrives,hovers,makesherchoiceandswoopsdown。Thetrickisdone:thepoorworker,withhertonguelollingfromhermouthinthedeath-struggle,iscarriedthroughtheairtotheundergroundden,whichisoftenaverylongwayfromthespotofthecapture。Thetricklingofearthyrefuse,onthebarebanks,orontheslopesoffootpaths,instantlyrevealsthedwellingsoftheravisher;and,asthePhilanthusalwaysworksinfairlypopulouscolonies,Iamable,bynotingthepositionofthecommunities,tomakesureoffruitfulexcavationsduringtheforcedinactivityofwinter。
Thesappingisalaborioustask,forthegalleriesruntoagreatdepth。
Favierwieldsthepickandspade;Ibreaktheclodswhichhebringsdownandopenthecells,whosecontents——cocoonsandremnantsofprovisions——I
atoncepourintoalittlescrewofpaper。Sometimes,whenthelarvaisnotdeveloped,thestackofBeesisintact;moreoftenthevictualshavebeenconsumed;butitisalwayspossibletotellthenumberofitemsprovided。
Theheads,abdomensandthoraxes,emptiedoftheirfleshysubstanceandreducedtothetoughouterskin,areeasilycounted。Ifthelarvahaschewedtheseovermuch,thewingsatleastareleft;thesearesaplessorganswhichthePhilanthusabsolutelyscorns。Theyarelikewisesparedbymoisture,putrefactionandtime,somuchsothatitisnomoredifficulttotakeaninventoryofacellseveralyearsoldthanoneofarecentcell。
Theessentialthingisnottooverlookanyofthesetinyrelicswhileplacingtheminthepaperbag,amidthethousandincidentsoftheexcavation。Therestoftheworkwillbedoneinthestudy,withtheaidofthelens,takingtheremainsheapbyheap;thewingswillbeseparatedfromthesurroundingrefuseandcountedinsetsoffour。Theresultwillgivetheamountoftheprovisions。Idonotrecommendthistasktoanyonewhoisnotendowedwithagoodstockofpatience,norabovealltoanyonewhodoesnotstartwiththeconvictionthatresultsofgreatinterestarecompatiblewithverymodestmeans。
Myinspectioncoversatotalofonehundredandthirty-sixcells,whicharedividedasinthetablebelow:
2cellseachcontaining1Bee52cellseachcontaining2Bees36cellseachcontaining3Bees36cellseachcontaining4Bees9cellseachcontaining5Bees1cellcontaining6Bees——
136
TheMantis-huntingTachytesconsumesitsheapofMantes,thehornyenvelopeincluded,withoutleavinganyremainsbutscantycrumbs,quiteinsufficienttoestablishthenumberofitemsprovided。Afterthemealiscompleted,anyinventoryoftherationsbecomesimpossible。Ithereforehaverecoursetothecellswhichstillcontaintheeggortheveryyounglarvaand,aboveall,tothosewhoseprovisionshavebeeninvadedbyatinyparasiticGnat,aTachina(Cf。“TheHuntingWasps“:chapters4and16——Translator\'sNote。),whichdrainsthegamewithoutcuttingitupandleavesthewholeskinintact。Twenty-fivelarders,puttothecount,givemethefollowingresult:
8cellseachcontaining3items5cellseachcontaining4items4cellseachcontaining6items3cellseachcontaining7items2cellseachcontaining8items1cellcontaining9items1cellcontaining12items1cellcontaining16items——
25
ThepredominantgameisthePrayingMantis,green;nextcomestheGreyMantis,ash-coloured。AfewEmpusaemakeupthetotal。Thespecimensvaryindimensionswithinfairlyelasticlimits:Imeasuresomewhichareathirdtoahalfinchlong,averagingtwo-thirdstooneinchlong,andsomewhicharetwo-fifths,averagingthreequarters。Iseeprettyplainlythattheirnumberincreasesinproportionastheirsizediminishes,asthoughtheTachyteswereseekingtomakeupforthesmallnessofthegamebyincreasingtheamount;nonethelessIfinditquiteimpossibletodetecttheleastequivalencebycombiningthetwofactorsofnumberandsize。Ifthehuntressreallyestimatestheprovisions,shedoessoveryroughly;herhouseholdaccountsarenotatallwellkept;eachheadofgame,largeorsmall,mustalwayscountasoneinhereyes。
Putonmyguard,Ilooktoseewhetherthehoney-gatheringBeeshaveadoubleservice,likethegame-huntingWasps\'。Iestimatetheamountofhoneyedpaste;Igaugethecupsintendedtocontainit。Inmanycasestheresultresemblesthefirstobtained:theabundanceofprovisionsvariesfromonecelltoanother。CertainOsmiae(O。cornutaandO。tricornis(Cf。
“Bramble-beesandOthers“:passim;and,inparticular,chapters3to5——
Translator\'sNote。))feedtheirlarvaeonaheapofpollen-dustmoistenedinthemiddlewithaverylittledisgorgedhoney。Oneoftheseheapsmaybethreeorfourtimesthesizeofsomeotherinthesamegroupofcells。IfI
detachfromitspebblethenestoftheMason-bee,theChalicodomaoftheWalls,Iseecellsoflargecapacity,sumptuouslyprovisioned;closebesidetheseIseeothers,oflesscapacity,withvictualsparsimoniouslyallotted。Thefactisgeneral;anditisrightthatweshouldaskourselvesthereasonforthesemarkeddifferencesintherelativequantityoffoodstuffsandfortheseunequalrations。
Iatlastbegantosuspectthatthisisfirstandforemostaquestionofsex。InmanyBeesandWasps,indeed,themaleandthefemalediffernotonlyincertaindetailsofinternalorexternalstructure——apointofviewwhichdoesnotaffectthepresentproblem——butalsoinlengthandbulk,whichdependinahighdegreeonthequantityoffood。
LetusconsiderinparticulartheBee-eatingPhilanthus。Comparedwiththefemale,themaleisamereabortion。Ifindthatheisonlyathirdtohalfthesizeoftheothersex,asfarasIcanjudgebysightalone。Toobtainexactlytherespectivequantitiesofsubstance,Ishouldneeddelicatebalances,capableofweighingdowntoamilligramme。Myclumsyvillager\'sscales,onwhichpotatoesmaybeweighedtowithinakilogrammeorso,donotpermitofthisprecision。Imustthereforerelyontheevidenceofmysightalone,evidence,forthatmatter,whichisamplysufficientinthepresentinstance。Comparedwithhismate,theMantis-huntingTachytesislikewiseapigmy。Wearequiteastonishedtoseehimpesteringhisgiantessonthethresholdoftheburrows。
Weobservedifferencesnolesspronouncedofsize——andconsequentlyofvolume,massandweight——inthetwosexesofmanyOsmiae。Thedifferencesarelessemphatic,butarestillonthesameside,intheCerceres,theStizi,theSpheges,theChalicodomaeandmanymore。Itisthereforetherulethatthemaleissmallerthanthefemale。Thereareofcoursesomeexceptions,thoughnotmany;andIamfarfromdenyingthem。IwillmentioncertainAnthidiawherethemaleisthelargerofthetwo。Nevertheless,inthegreatmajorityofcasesthefemalehastheadvantage。
Andthisisasitshouldbe。Itisthemother,themotheralone,wholaboriouslydigsundergroundgalleriesandchambers,kneadstheplasterforcoatingthecells,buildsthedwelling-houseofcementandbitsofgrit,boresthewoodanddividestheburrowintostoreys,cutsthedisksofleafwhichwillbejoinedtogethertoformhoney-pots,worksuptheresingatheredindropsfromthewoundsinthepine-treestobuildceilingsintheemptyspiralofaSnail-shell,huntstheprey,paralysesitanddragsitindoors,gathersthepollen-dust,preparesthehoneyinhercrop,storesandmixesthepaste。Thisseverelabour,soimperiousandsoactive,inwhichtheinsect\'swholelifeisspent,manifestlydemandsabodilystrengthwhichwouldbequiteuselesstothemale,theamoroustrifler。
Thus,asageneralrule,intheinsectswhichcarryonanindustrythefemaleisthestrongersex。
Doesthispre-eminenceimplymoreabundantprovisionsduringthelarvalstage,whentheinsectisacquiringthephysicalgrowthwhichitwillnotexceedinitsfuturedevelopment?Simplereflectionsuppliestheanswer:
yes,theaggregategrowthhasitsequivalentintheaggregateprovisions。
ThoughsoslightacreatureasthemalePhilanthusfindsarationoftwoBeessufficientforhisneeds,thefemale,twiceorthriceasbulky,willconsumethreetosixatleast。IfthemaleTachytesrequiresthreeMantes,hisconsort\'smealwilldemandabatchofsomethingliketen。Withhercomparativecorpulence,thefemaleOsmiawillneedaheapofpastetwiceorthriceasgreatasthatofherbrother,themale。Allthisisobvious;theanimalcannotmakemuchoutoflittle。
Despitethisevidence,Iwasanxioustoenquirewhethertherealitycorrespondedwiththeprevisionsofthemostelementarylogic。Instancesarenotunknowninwhichthemostsagaciousdeductionshavebeenfoundtodisagreewiththefacts。Duringthelastfewyears,therefore,Ihaveprofitedbymywinterleisuretocollect,fromspotsnotedasfavourableduringtheworking-season,afewhandfulsofcocoonsofvariousDigger-
wasps,notablyoftheBee-eatingPhilanthus,whohasjustfurnisheduswithaninventoryofprovisions。Surroundingthesecocoonsandthrustagainstthewallofthecellweretheremnantsofthevictuals——wings,corselets,heads,wing-cases——acountofwhichenabledmetodeterminehowmanyheadofgamehadbeenprovidedforthelarva,nowenclosedinitssilkenabode。
Ithusobtainedthecorrectlistofprovisionsforeachofthehuntress\'
cocoons。Ontheotherhand,Iestimatedthequantitiesofhoney,orratherIgaugedthereceptacles,thecells,whosecapacityisproportionatetothemassoftheprovisionsstored。Aftermakingthesepreparations,registeringthecells,cocoonsandrationsandputtingallmyfiguresinorder,Ihadonlytowaitforthehatching-seasontodeterminethesex。
Well,Ifoundthatlogicandexperimentwereinperfectagreement。ThePhilanthus-cocoonswithtwoBeesgavememales,alwaysmales;thosewithalargerrationgavemefemales。FromtheTachytes-cocoonswithdoubleortreblethatrationIobtainedfemales。WhenfeduponfourorfiveNut-
weevils,theSandCerceriswasamale;whenfeduponeightorten,afemale。Inshort,abundantprovisionsandspaciouscellsyieldfemales;
scantyprovisionsandnarrowcellsyieldmales。ThisisalawuponwhichI
mayhenceforthrely。
Atthestagewhichwehavenowreachedaquestionarises,aquestionofmajorimportance,touchingthemostnebulousaspectofembryogeny。HowisitthatthelarvaofthePhilanthus,totakeaparticularcase,receivesthreetofiveBeesfromitsmotherwhenitistobecomeafemaleandnotmorethantwowhenitistobecomeamale?Herethevariousheadofgameareidenticalinsize,inflavour,innutritiveproperties。Thefood-valueispreciselyinproportiontothenumberofitemssupplied,ahelpfuldetailwhicheliminatestheuncertaintieswhereinwemightbeleftbytheprovisionofgameofdifferentspeciesandvaryingsizes。Howisit,then,thatahostofBeesandWasps,ofhoney-gatherersaswellashuntresses,storealargerorsmallerquantityofvictualsintheircellsaccordingasthenurselingsaretobecomefemalesormales?
Theprovisionsarestoredbeforetheeggsarelaid;andtheseprovisionsaremeasuredbytheneedsofthesexofaneggstillinsidethemother\'sbody。Iftheegg-layingweretoprecedetherationing,whichoccasionallytakesplace,aswiththeOdyneri(Cf。“TheHuntingWasps“:chapters2and8——Translator\'sNote。),forexample,wemightimaginethatthegravidmotherenquiresintothesexoftheegg,recognizesitandstacksvictualsaccordingly。But,whetherdestinedtobecomeamaleorafemale,theeggisalwaysthesame;thedifferences——andIhavenodoubtthattherearedifferences——areinthedomainoftheinfinitelysubtle,themysterious,imperceptibleeventothemostpractisedembryogenist。Whatcanapoorinsectsee——intheabsolutedarknessofitsburrow,moreover——wheresciencearmedwithopticalinstrumentshasnotyetsucceededinseeinganything?
Andbesides,evenwereitmorediscerningthanweareinthesegeneticobscurities,itsvisualdiscernmentwouldhavenothingwhereupontopractice。AsIhavesaid,theeggislaidonlywhenthecorrespondingprovisionsarestored。Themealispreparedbeforethelarvawhichistoeatithascomeintotheworld。Thesupplyisgenerouslycalculatedbytheneedsofthecomingcreature;thedining-roomisbuiltlargeorsmalltocontainagiantoradwarfstillgerminatingintheovarianducts。Themother,therefore,knowsthesexofhereggbeforehand。
Astrangeconclusion,whichplayshavocwithourcurrentnotions!Thelogicofthefactsleadsustoitdirectly。Andyetitseemssoabsurdthat,beforeacceptingit,weseektoescapethepredicamentbyanotherabsurdity。Wewonderwhetherthequantityoffoodmaynotdecidethefateoftheegg,originallysexless。Givenmorefoodandmoreroom,theeggwouldbecomeafemale;givenlessfoodandlessroom,itwouldbecomeamale。Themother,obeyingherinstincts,wouldstoremorefoodinthiscaseandlessinthat;shewouldbuildnowalargeandnowasmallcell;andthefutureoftheeggwouldbedeterminedbytheconditionsoffoodandshelter。
Letusmakeeverytest,everyexperiment,downtotheabsurd:thecrudeabsurdityofthemomenthassometimesprovedtobethetruthofthemorrow。
Besides,thewell-knownstoryoftheHive-beeshouldmakeuswaryofrejectingparadoxicalsuppositions。Isitnotbyincreasingthesizeofthecell,bymodifyingthequalityandquantityofthefood,thatthepopulationofahivetransformsaworkerlarvaintoafemaleorroyallarva?Itistruethatthesexremainsthesame,sincetheworkersareonlyincompletelydevelopedfemales。Thechangeisnonethelessmiraculous,somuchsothatitisalmostlawfultoenquirewhetherthetransformationmaynotgofurther,turningamale,thatpoorabortion,intoasturdyfemalebymeansofaplentifuldiet。Letusthereforeresorttoexperiment。
IhaveathandsomelongbitsofreedinthehollowofwhichanOsmia,theThree-hornedOsmia,hasstackedhercells,boundedbyearthenpartitions。I
haverelatedelsewhere(Cf。“Bramble-beesandOthers“:chapters2to5——
Translator\'sNote。)howIobtainasmanyofthesenestsasIcouldwishfor。Whenthereedissplitlengthwise,thecellscomeintoview,togetherwiththeirprovisions,theegglyingonthepaste,oreventhebuddinglarva。Observationsmultipliedadnauseamhavetaughtmewheretofindthemalesandwherethefemalesinthisapiary。Themalesoccupythefore-partofthereed,theendnexttotheopening;thefemalesareatthebottom,nexttotheknotwhichservesasanaturalstoppertothechannel。Fortherest,thequantityoftheprovisionsinitselfpointstothesex:forthefemalesitistwiceorthriceasgreatasforthemales。
Inthescantily-providedcells,Idoubleortrebletherationwithfoodtakenfromothercells;inthecellswhichareplentifullysupplied,I
reducetheportiontoahalforathird。Controlsareleft:thatistosay,somecellsremainuntouched,withtheirprovisionsasIfoundthem,bothinthepartwhichisabundantlyprovidedandinthatwhichismoremeagrelyrationed。Thetwohalvesofthereedarethenrestoredtotheiroriginalpositionandfirmlyboundwithafewturnsofwire。Weshallsee,whenthetimecomes,whetherthesechangesincreasingordecreasingthevictualshavedeterminedthesex。
Hereistheresult:thecellswhichatfirstweresparinglyprovided,butwhosesuppliesweredoubledortrebledbymyartifice,containmales,asforetoldbytheoriginalamountofvictuals。ThesurpluswhichIaddedhasnotcompletelydisappeared,farfromit:thelarvahashadmorethanitneededforitsevolutionasamale;and,beingunabletoconsumethewholeofitscopiousprovisions,ithasspunitscocooninthemidstoftheremainingpollen-dust。Thesemales,sorichlysupplied,areofhandsomebutnotexaggeratedproportions;youcanseethattheadditionalfoodhasprofitedthemtosomesmallextent。
Thecellswithabundantprovisions,reducedtoahalforathirdbymyintervention,containcocoonsassmallasthemalecocoons,pale,translucentandlimp,whereasthenormalcocoonsaredark-brown,opaqueandfirmtothetouch。These,weperceiveatonce,aretheworkofstarved,anaemicweavers,who,failingtosatisfytheirappetiteandhavingeatenthelastgrainofpollen,have,beforedying,donetheirbestwiththeirpoorlittledropofsilk。Thosecocoonswhichcorrespondwiththesmallestallowanceoffoodcontainonlyadeadandshrivelledlarva;others,inwhosecasetheprovisionswerelessmarkedlydecreased,containfemalesintheadultform,butofverydiminutivesize,comparablewiththatofthemales,orevensmaller。AsforthecontrolswhichIwascarefultoleave,theyconfirmthefactthatIhadmalesinthepartneartheorificeofthereedandfemalesinthepartneartheknotclosingthechannel。
Isthisenoughtodisposeoftheveryimprobablesuppositionthatthedeterminationofthesexdependsonthequantityoffood?Strictlyspeaking,thereisstillonedooropentodoubt。Itmaybesaidthatexperiment,withitsartifices,doesnotsucceedinrealizingthedelicatenaturalconditions。Tomakeshortworkofallobjections,Icannotdobetterthanhaverecoursetofactsinwhichtheexperimenter\'shandhasnotintervened。Theparasiteswillsupplyuswiththesefacts;theywillshowushowalienthequantityandeventhequalityofthefoodarefromeitherspecificorsexualcharacters。Thesubjectofenquirythusbecomesdouble,insteadofsingleasitwaswhenIplunderedonecellinmysplitreedstoenrichanother。Letusfollowthisdoublecurrentforalittlewhile。
AnAmmophila,theSilkyAmmophila(Cf。“TheHuntingWasps“:chapter13——
Translator\'sNote。),whichfeedsonLoopercaterpillars(KnownalsoasMeasuring-worms,Inchworms,SpanwormsandSurveyors:thecaterpillarsoftheGeometridMoths——Translator\'sNote。),hasjustbeenrearedinmyrefectoryonSpiders。Repletetotheregulationpoint,itspinsitscocoon。
Whatwillemergefromthis?Ifthereaderexpectstoseeanymodifications,causedbyadietwhichthespecies,lefttoitself,hadnevereffected,lethimbeundeceivedandthatquickly。TheAmmophilafedonSpidersispreciselythesameastheAmmophilafedoncaterpillars,justasmanfedonriceisthesameasmanfedonwheat。InvainIpassmylensovertheproductofmyart:Icannotdistinguishitfromthenaturalproduct;andI
defythemostmeticulousentomologisttoperceiveanydifferencebetweenthetwo。Itisthesamewithmyotherboarderswhohavehadtheirdietaltered。
Iseetheobjectioncoming。Thedifferencesmaybeinappreciable,formyexperimentstouchonlyafirstrungoftheladder。Whatwouldhappeniftheladderwereprolonged,iftheoffspringoftheAmmophilafedonSpidersweregiventhesamefoodgenerationaftergeneration?Thesedifferences,atfirstimperceptible,mightbecomeaccentuateduntiltheygrewintodistinctspecificcharacters;thehabitsandinstinctsmightalsochange;andintheendthecaterpillar-huntressmightbecomeaSpider-huntress,withashapeofherown。Aspecieswouldbecreated,for,amongthefactorsatworkinthetransformationofanimals,themostimportantofallisincontestablythetypeoffood,thenatureofthethingwherewiththeanimalbuildsitself。AllthisismuchmoreimportantthanthetrivialitieswhichDarwinreliesupon。
Tocreateaspeciesismagnificentintheory,sothatwefindourselvesregrettingthattheexperimenterisnotabletocontinuetheattempt。But,oncetheAmmophilahasflownoutofthelaboratorytoslakeherthirstattheflowersintheneighbourhood,justtotrytofindheragainandinducehertoentrustyouwithhereggs,whichyouwouldrearintherefectory,toincreasethetasteforSpidersfromgenerationtogeneration!Merelytodreamofitweremadness。Shallwe,inourhelplessness,admitourselvesbeatenbytheevolutionaryeffectsofdiet?Notabitofit!Oneexperiment——andyoucouldnotwishforamoredecisive——iscontinuallyinprogress,apartfromallartifices,onanenormousscale。Itisbroughttoournoticebytheparasites。
Theymust,wearetold,haveacquiredthehabitoflivingonothersinordertosavethemselvesworkandtoleadaneasierlife。Thepoorwretcheshavemadeasorryblunder。Theirlifeisofthehardest。Ifafewestablishthemselvescomfortably,dearthanddirefamineawaitmostoftherest。
Therearesome——lookatcertainoftheOil-beetles——exposedtosomanychancesofdestructionthat,tosaveone,theyareobligedtoprocreateathousand。Theyseldomenjoyafreemeal。Somestrayintothehousesofhostswhosevictualsdonotsuitthem;othersfindonlyarationquiteinsufficientfortheirneeds;others——andtheseareverynumerous——findnothingatall。Whatmisadventures,whatdisappointmentsdotheseneedycreaturessuffer,unaccustomedastheyaretowork!Letmerelatesomeoftheirmisfortunes,gleanedatrandom。
TheGirdledDioxys(D。cincta)lovestheamplehoney-storesoftheChalicodomaofthePebbles。Thereshefindsabundantfood,soabundantthatshecannoteatitall。Ihavealreadypassedcensureonthiswaste。(Cf。
“TheMason-bees“:chapter10——Translator\'sNote。)NowalittleOsmia(O。
cyanoxantha,Perez)makeshernestintheMason\'sdesertedcells;andthisBee,avictimofherill-omeneddwelling,alsoharbourstheDioxys。Thisisamanifesterrorontheparasite\'spart。ThenestoftheChalicodoma,thehemisphereofmortaronitspebble,iswhatsheislookingfor,toconfidehereggstoit。Butthenestisnowoccupiedbyastranger,bytheOsmia,acircumstanceunknowntotheDioxys,whocomesstealinguptolayheregginthemother\'sabsence。Thedomeisfamiliartoher。Shecouldnotknowitbetterifshehadbuiltitherself。Hereshewasborn;hereiswhatherfamilywants。Moreover,thereisnothingtoarousehersuspicions:theoutsideofthehomehasnotchangeditsappearanceinanyrespect;thestopperofgravelandgreenputty,whichlaterwillformaviolentcontrastwithitswhitefront,isnotyetconstructed。Shegoesinandseesaheapofhoney。ToherthinkingthiscanbenothingbuttheChalicodoma\'sportion。Weourselveswouldbebeguiled,intheOsmia\'sabsence。Shelayshereggsinthisdeceptivecell。
Hermistake,whichiseasytounderstand,doesnotinanywaydetractfromhergreattalentsasaparasite,butitisaseriousmatterforthefuturelarva。TheOsmia,infact,inviewofhersmalldimensions,collectsbutaveryscantystoreoffood:alittleloafofpollenandhoney,hardlythesizeofanaveragepea。SucharationisinsufficientfortheDioxys。I
havedescribedherasawasteroffoodwhenherlarvaisestablished,accordingtocustom,inthecelloftheMason-bee。Thisdescriptionnolongerapplies;notintheveryleast。InadvertentlystrayingtotheOsmia\'stable,thelarvahasnoexcuseforturningupitsnose;itdoesnotleavepartofthefoodtogobad;iteatsupthelotwithouthavinghadenough。
Thisfamine-strickenrefectorycangiveusnothingbutanabortion。Asamatteroffact,theDioxyssubjectedtothisniggardlytestdoesnotdie,fortheparasitemusthaveatoughconstitutiontoenableittofacethedisastroushazardswhichlieinwaitforit;butitattainsbarelyhalfitsordinarydimensions,whichmeansone-eighthofitsnormalbulk。Toseeitthusdiminished,wearesurprisedatitstenaciousvitality,whichenablesittoreachtheadultforminspiteoftheextremedeficiencyoffood。
Meanwhile,thisadultisstilltheDioxys;thereisnochangeofanykindinhershapeorcolouring。Moreover,thetwosexesarerepresented;thisfamilyofpigmieshasitsmalesandfemales。DearthandthefarinaceousmessintheOsmia\'scellhashadnomoreinfluenceoverspeciesorsexthanabundanceandflowinghoneyintheChalicodoma\'shome。
ThesamemaybesaidoftheSpottedSapyga(S。punctata(AparasiticWasp。
Cf。“TheMason-bees“:chapters9and10——Translator\'sNote。)),which,aparasiteoftheThree-prongedOsmia,adenizenofthebramble,andoftheGoldenOsmia,anoccupantofemptySnail-shells,straysintothehouseoftheTinyOsmia(O。parvula(Thisbeemakesherhomeinthebrambles。Cf。
“Bramble-dwellersandOthers“:chapters2and3——Translator\'sNote。)),where,forlackofsufficientfood,itdoesnotattainhalfitsnormalsize。
ALeucopsis(Cf。“TheMason-bees“:chapter11——Translator\'sNote。)insertshereggsthroughthecementwallofourthreeChalicodomae。Iknowherundertwonames。WhenshecomesfromtheChalicodomaofthePebblesorWalls,whoseopulentlarvasaturatesherwithfood,shedeservesbyherlargesizethenameofLeucopsisgigas,whichFabriciusbestowsuponher;
whenshecomesfromtheChalicodomaoftheSheds,shedeservesnomorethanthenameofL。grandis,whichisallthatKluggrantsher。Withasmallerration“thegiant“istosomedegreediminishedandbecomesnomorethan“thelarge。“WhenshecomesfromtheChalicodomaoftheShrubs,sheissmallerstill;and,ifsomenomenclatorweretoseektodescribeher,shewouldnolongerdeservetobecalledmorethanmiddling。Fromdimension2
shehasdescendedtodimension1withoutceasingtobethesameinsect,despitethechangeofdiet;andatthesametimebothsexesarepresentinthethreenurselings,despitethevariationinthequantityofvictuals。
IobtainAnthraxsinuata(“TheMason-bees“:chapters8,10and11——
Translator\'sNote。)fromvariousbees\'nests。WhensheissuesfromthecocoonsoftheThree-hornedOsmia,especiallythefemalecocoons,sheattainsthegreatestdevelopmentthatIknowof。WhensheissuesfromthecocoonsoftheBlueOsmia(O。cyanea,KIRB。),sheissometimeshardlyone-
thirdthelengthwhichtheotherOsmiagivesher。Andwestillhavethetwosexes——thatgoeswithoutsaying——andstillidenticallythesamespecies。
TwoAnthidia,workinginresin,A。septemdentatum,LATR。,andA。
bellicosum,LEP。(FortheseResin-bees,cf。“Bramble-beesandOthers“:
chapter10——Translator\'sNote。),establishtheirdomicileinoldSnail-
shells。ThesecondharbourstheBurntZonitis(Z。proeusta(Cf。“TheGlow-
wormandOtherBeetles“:chapter6——Translator\'sNote。))。AmplynourishedthisMeloethenacquireshernormalsize,thesizeinwhichsheusuallyfiguresinthecollections。AlikeprosperityawaitsherwhensheusurpstheprovisionsofMegachilesericans。(ForthisBee,theSilkyLeaf-cutter,cf。“Bramble-beesandOthers“:chapter8——Translator\'sNote。)ButtheimprudentcreaturesometimesallowsitselftobecarriedawaytothemeagretableofthesmallestofourAnthidia(A。scapulare,LATR。(ACotton-bee,cf。idem:chapter9——Translator\'sNote。)),whomakeshernestsindrybramble-stems。Thescantyfaremakesawretcheddwarfoftheoffspringbelongingtoeithersex,withoutdeprivingthemofanyoftheirracialfeatures。WestillseetheBurntZonitis,withthedistinctivesignofthespecies:thesingedpatchatthetipofthewing-cases。
AndtheotherMeloidae——Cantharides,Cerocomae,Mylabres(FortheseBlister-beetlesorOil-beetles,cf。“TheGlow-wormandOtherBeetles“:
chapter6——Translator\'sNote。)——towhatinequalitiesofsizearetheynotsubject,irrespectiveofsex!Therearesome——andtheyarenumerous——whosedimensionsfalltoahalf,athird,aquarteroftheregulardimensions。
Amongthesedwarfs,thesemisbegottenones,thesevictimsofatrophy,therearefemalesaswellasmales;andtheirsmallnessbynomeanscoolstheiramorousardour。Theseneedycreatures,Irepeat,haveahardlifeofit。
Whencedotheycome,thesediminutiveBeetles,ifnotfromdining-roomsinsufficientlysuppliedfortheirneeds?Theirparasiticalhabitsexposethemtoharshvicissitudes。Nomatter:indearthaswellasinabundancethetwosexesappearandthespecificfeaturesremainunchanged。
Itisunnecessarytolingerlongeroverthissubject。Thedemonstrationiscompleted。Theparasitestellusthatchangesinthequantityandqualityoffooddonotleadtoanytransformationofspecies。FeduponthelarvaoftheThree-hornedOsmiaoroftheBlueOsmia,Anthraxsinuata,whetherofhandsomeproportionsoradwarf,isstillAnthraxsinuata;fedupontheallowanceoftheAnthidiumoftheemptySnail-shells,theAnthidiumofthebrambles,theMegachileordoubtlessmanyothers,theBurntZonitisisstilltheBurntZonitis。Yetvariationofdietoughttobeaverypotentialfactorintheproblemofprogresstowardsanotherform。Isnottheworldoflivingcreaturesruledbythestomach?Andthevalueofthisfactorisunity,changingnothingintheproduct。
Thesameparasitestellus——andthisisthechiefobjectofmydigression——
thatexcessordeficiencyofnutrimentdoesnotdeterminethesex。Soweareoncemoreconfrontedwiththestrangeproposition,whichisnowmorepositivethanever,thattheinsectwhichamassesprovisionsinproportiontotheneedsoftheeggabouttobelaidknowsbeforehandwhatthesexofthiseggwillbe。Perhapstherealityisevenmoreparadoxicalstill。I
shallreturntothesubjectafterdiscussingtheOsmiae,whoareveryweightywitnessesinthisgraveaffair。(Cf。“Bramble-beesandOthers“:
chapters3to5。Thestudentisrecommendedtoreadthesethreechaptersinconjunctionwiththepresentchapter,towhichtheyformasequel,withthatontheOsmiae(chapter2oftheabovevolume)intervening——
Translator\'sNote。)
CHAPTER10。THEBEE-EATINGPHILANTHUS。
TomeetamongtheWasps,thoseeagerloversofflowers,aspeciesthatgoeshuntingmoreorlessonitsownaccountiscertainlyanotableevent。Thatthelarderofthegrubshouldbeprovidedwithpreyisnaturalenough;butthattheprovider,whosedietishoney,shouldherselfmakeuseofthecaptivesisanythingbuteasytounderstand。Wearequiteastonishedtoseeanectar-drinkerbecomeablood-drinker。Butourastonishmentceasesifweconsiderthingsmoreclosely。Thedoublemethodoffeedingismoreapparentthanreal:thecropwhichfillsitselfwithsugaryliquiddoesnotgorgeitselfwithgame。TheOdynerus,whendiggingintothebodyofherprey,doesnottouchtheflesh,afareabsolutelyscornedascontrarytohertastes;shesatisfiesherselfwithlappingupthedefensivedropwhichthegrub(TheLarvaofChrysomelapopuli,thePoplarLeaf-beetle——Translator\'sNote。)distilsattheendofitsintestine。Thisfluidnodoubtrepresentstohersomehighly-flavouredbeveragewithwhichsheseasonsfromtimetotimethestapledietfetchedfromthedrinking-baroftheflowers,someappetizingcondimentorperhaps——whoknows?——somesubstituteforhoney。
Thoughthequalitiesofthedelicacyescapeme,IatleastperceivethattheOdynerusdoesnotcovetanythingelse。Onceitsjarisemptied,thelarvaisflungasideasworthlessoffal,acertainsignofanon-
carnivorousappetite。Undertheseconditions,thepersecutoroftheChrysomelaceasestosurpriseusbyindulginginthecryingabuseofadoublediet。
Weevenbegintowonderwhetherotherspeciesmaynotbeinclinedtoderiveadirectadvantagefromthehuntingimposeduponthemforthemaintenanceofthefamily。TheOdynerus\'methodofwork,thesplittingopenoftheanalstill-room,istoofarremovedfromtheobviousproceduretohavemanyimitators;itisasecondarydetailandimpracticablewithadifferentkindofgame。Butthereissuretobeacertainvarietyinthedirectmeansofutilizingthecapture。Why,forinstance,whenthevictimparalysedbythestingcontainsadeliciousbrothinsomepartofitsstomach,shouldthehuntressscrupletoviolateherdyingpreyandforceittodisgorgewithoutinjuringthequalityoftheprovisions?Theremustbethosewhorobthedead,attractednotbythefleshbutbytheexquisitecontentsofthecrop。
Inpointoffact,thereare;andtheyareevennumerous。WemaymentioninthefirstranktheWaspthathuntsHive-bees,theBee-eatingPhilanthus(P。
apivorus,LATR。)。Ilongsuspectedherofperpetratingtheseactsofbrigandageonherownbehalf,havingoftensurprisedhergluttonouslylickingtheBee\'shoney-smearedmouth;Ihadaninklingthatshedidnotalwayshuntsolelyforthebenefitofherlarvae。Thesuspiciondeservedtobeconfirmedbyexperiment。Also,Iwasengagedinanotherinvestigation,whichmighteasilybeconductedsimultaneouslywiththeonesuggested:I
wantedtostudy,withalltheleisureofworkdoneathome,theoperating-
methodsemployedbythedifferentHuntingWasps。Ithereforemadeuse,forthePhilanthus,oftheprocessofexperimentingunderglasswhichIroughlyoutlinedwhenspeakingoftheOdynerus。ItwaseventheBee-huntresswhogavememyfirstdatainthisdirection。SherespondedtomywisheswithsuchzealthatIbelievedmyselftopossessanunequalledmeansofobservingagainandagain,eventoexcess,whatissodifficulttoachieveontheactualspot。Alas,thefirst-fruitsofmyacquaintancewiththePhilanthuspromisedmemorethanthefutureheldinstoreforme!Butwewillnotanticipate;andwewillplacethehuntressandhergametogetherunderthebell-glass。IrecommendthisexperimenttowhoeverwouldwishtoseewithwhatperfectionintheartofattackanddefenceaHuntingWaspwieldsthestiletto。Thereisnouncertaintyhereastotheresult,thereisnolongwait:themomentwhenshecatchessightofthepreyinanattitudefavourabletoherdesigns,thebanditrushesforwardandkills。I
willdescribehowthingshappen。
Iplaceunderthebell-glassaPhilanthusandtwoorthreeHive-bees。Theprisonersclimbtheglasswall,towardsthelight;theygoup,comedownagainandtrytogetout;theverticalpolishedsurfaceistothemapracticablefloor。Theysoonquietdown;andthespoilerbeginstonoticehersurroundings。Theantennaearepointedforwards,enquiringly;thehind-
legsaredrawnupwithalittlequiverofgreedinthetarsi;theheadturnstorightandleftandfollowstheevolutionsoftheBeesagainsttheglass。Themiscreant\'sposturenowbecomesastrikingpieceofacting:youcanreadinitthefiercelongingsofthecreaturelyinginambush,thecraftywaitingforthemomenttocommitthecrime。Thechoiceismade:thePhilanthuspouncesonherprey。
Turnbyturntumblingoverandtumbled,thetwoinsectsrollupontheground。Thetumultsoonabates;andthemurderesspreparestostranglehercapture。Iseeheradopttwomethods。Inthefirst,whichismoreusualthantheother,theBeeislyingonherback;andthePhilanthus,bellytobellywithher,gripsherwithhersixlegswhilesnappingatherneckwithhermandibles。Theabdomenisnowcurvedforwardfrombehind,alongtheprostratevictim,feelswithitstip,gropesaboutalittleandendsbyreachingtheunderpartoftheneck。Thestingenters,lingersforamomentinthewound;andallisover。Withoutreleasingherprey,whichisstilltightlyclasped,themurderessrestoresherabdomentoitsnormalpositionandkeepsitpressedagainsttheBee\'s。
Inthesecondmethod,thePhilanthusoperatesstanding。Restingonherhind-legsandonthetipsofherunfurledwings,sheproudlyoccupiesanerectattitude,withtheBeeheldfacingherbetweenherfourfrontlegs。
Togivethepoorthingapositionsuitedtoreceivethedagger-stroke,sheturnsherroundandbackagainwiththeroughclumsinessofachildhandlingitsdoll。Herposeismagnificenttolookat。Solidlyplantedonhersustainingtripod,thetwohindertarsiandthetipsofthewings,sheatlastcrooksherabdomenupwardsandagainstingstheBeeunderthechin。
TheoriginalityofthePhilanthus\'postureatthemomentofthemurdersurpassestheanythingthatIhavehithertoseen。
Thedesireforknowledgeinnaturalhistoryhasitscruelside。Tolearnpreciselythepointattackedbythestingandtomakemyselfthoroughlyacquaintedwiththehorribletalentofthemurderess,IhaveinvestigatedmoreassassinationsunderglassthanIwoulddaretoconfess。Withoutasingleexception,IhavealwaysseentheBeestunginthethroat。Inthepreparationsforthefinalblow,thetipoftheabdomenmaywellcometorestonthisorthatpointofthethoraxorabdomen;butitdoesnotstopatanyofthese,noristhestingunsheathed,ascanreadilybeascertained。Indeed,oncethecontestisopened,thePhilanthusbecomessoentirelyabsorbedinheroperationthatIcanremovethecoverandfolloweveryvicissitudeofthetragedywithmypocket-lens。
Afterrecognizingtheinvariablepositionofthewound,Ibendbackandopenthearticulationofthehead。IseeundertheBee\'schinawhitespot,measuringhardlyatwenty-fifthofaninchsquare,wherethehornyintegumentsarelackingandthedelicateskinisshownuncovered。Itishere,alwayshere,inthistinydefectinthearmour,thatthestingenters。Whyisthisspotstabbedratherthananother?Canitbetheonlyvulnerablepoint,whichwouldnecessarilydeterminethethrustofthelancet?Shouldanyoneentertainsopettyathought,Iadvisehimtoopenthearticulationofthecorselet,behindthefirstpairoflegs。HewillthereseewhatIsee:thebareskin,quiteasfineasundertheneck,butcoveringamuchlargersurface。Thehornybreast-plateoffersnowiderbreach。IfthePhilanthuswereguidedinheroperationsolelybythequestionofvulnerability,itisherecertainlythatsheoughttostrike,insteadofpersistentlyseekingthenarrowslitintheneck。Theweaponwouldnotneedtohesitateandgrope;itwouldobtainadmissionintothetissuesoff-hand。No,thestrokeofthelancetisnotforceduponitmechanically:theassassinscornsthelargedefectinthecorseletandpreferstheplaceunderthechin,foreminentlylogicalreasonswhichwewillnowattempttounravel。
ImmediatelyaftertheoperationItaketheBeefromthePhilanthus。Whatstrikesmeisthesuddeninertiaoftheantennaeandthemouth-parts,organswhichinthevictimsofmostoftheHuntingWaspscontinuetomoveforsolongatime。ThereareherenotanyofthesignsoflifetowhichI
havebeenaccustomedinmyoldstudiesofinsectparalysis:theantennarythreadswavingslowlytoandfro,thepalpiquivering,themandiblesopeningandclosingfordays,weeksandmonthsonend。Atmost,thetarsitrembleforaminuteortwo;thatconstitutesthewholedeath-struggle。
Completeimmobilityensues。Theinferencedrawnfromthissuddeninertiaisinevitable:theWasphasstabbedthecervicalganglia。Hencetheimmediatecessationofmovementinalltheorgansofthehead;hencetherealinsteadoftheapparentdeathoftheBee。ThePhilanthusisabutcherandnotaparalyser。
Thisisonestepgained。Themurderesschoosestheunderpartofthechinasthepointattackedinordertostriketheprincipalnerve-centres,thecephalicganglia,andthustodoawaywithlifeatoneblow。Whenthisvitalseatispoisonedbythetoxin,deathisinstantaneous。HadthePhilanthus\'objectbeensimplytoeffectparalysis,thesuppressionoflocomotormovements,shewouldhavedrivenherweaponintotheflawinthecorselet,astheCerceresdowiththeWeevils,whoaremuchmorepowerfullyarmouredthantheBee。Butherintentionistokilloutright,asweshallseepresently;shewantsacorpse,notaparalyticpatient。Thisbeingso,wemustagreethatheroperating-methodissupremelywell-inspired:ourhumanmurdererscouldachievenothingmorethoroughorimmediate。
Wemustalsoagreethatherattitudewhenattacking,anattitudeverydifferentfromthatoftheparalysers,isinfallibleinitsdeath-dealingefficacy。Whethershedeliverherthrustlyingonthegroundorstandingerect,sheholdstheBeeinfrontofher,breasttobreast,headtohead。
Inthispostureallthatsheneeddoistocurveherabdomeninordertoreachthegapintheneckandplungethestingwithanupwardslantintohercaptive\'shead。Supposethetwoinsectstobegrippingeachotherinthereverseattitude,imaginethedirktoslantslightlyintheoppositedirection;theresultswouldbeabsolutelydifferentandthesting,drivendownwards,wouldpiercethefirstthoracicganglionandproducemerelypartialparalysis。Whatskill,tosacrificeawretchedBee!Inwhatfencing-schoolwastheslayertaughtherterribleupwardblowunderthechin?
Ifshelearntit,howisitthathervictim,suchapastmistressinarchitecture,suchanadeptinsocialisticpolity,hassofarlearntnocorrespondingtricktoserveinherowndefence?Sheisaspowerfulasherexecutioner;liketheother,shecarriesarapier,anevenmoreformidableoneandmorepainful,atleasttomyfingers。ForcenturiesandcenturiesthePhilanthushasbeenstoringherawayinhercellars;andthepoorinnocentmeeklysubmits,withoutbeingtaughtbytheannualexterminationofherracehowtodeliverherselffromtheaggressorbyawell-aimedthrust。Idespairofeverunderstandinghowtheassailanthasacquiredhertalentforinflictingsuddendeath,whentheassailed,whoisbetter-armedandquiteasstrong,wieldsherdaggeranyhowandthereforeineffectively。