Iftheonehaslearntbyprolongedpracticeinattack,theothershouldalsohavelearntbyprolongedpracticeindefence,forattackanddefencepossessalikemeritinthefightforlife。Amongthetheoristsoftheday,isthereoneclear-sightedenoughtosolvetheriddleforus?
Ifso,Iwilltaketheopportunityofputtingtohimasecondproblemthatpuzzlesme:thecarelessness,nay,more,thestupidityoftheBeeinthepresenceofthePhilanthus。Youwouldbeinclinedtothinkthatthevictimofpersecution,learninggraduallyfromthemisfortunessufferedbyherfamily,wouldshowdistressattheravisher\'sapproachandatleastattempttoescape。InmycagesIseenothingofthesort。Oncethefirstexcitementduetoincarcerationunderthebell-glassorthewire-gauzecoverhaspassed,theBeeseemshardlytotroubleaboutherformidableneighbour。I
seeonesidebysidewiththePhilanthusonthesamehoneyedthistle-head:
assassinandfuturevictimaredrinkingfromthesameflask。Iseesomeonewhocomesheedlesslytoenquirewhothatstrangercanbe,crouchinginwaitonthetable。Whenthespoilermakesherrush,itisusuallyataBeewhomeetsherhalf-way,and,sotospeak,flingsherselfintoherclutches,eitherthoughtlesslyoroutofcuriosity。Thereisnowildterror,nosignofanxiety,notendencytomakeoff。Howcomesitthattheexperienceoftheages,thatexperiencewhich,wearetold,teachestheanimalsomanythings,hasnottaughttheBeethefirstelementofapiarianwisdom:adeep-seatedhorrorofthePhilanthus?Canthepoorwretchtakecomfortbyrelyingonhertrustydagger?Butsheyieldstononeinherignoranceoffencing;shestabswithoutmethod,atrandom。However,letuswatchheratthesuprememomentofthekilling。
Whentheravishermakesplaywithhersting,theBeedoesthesamewithhersandfuriously。Iseetheneedlenowmovingthiswayorthatwayinspace,nowslipping,violentlycurved,alongthemurderess\'convexsurface。
Thesesword-thrustshavenoseriousresults。Themannerinwhichthetwocombatantsareatgripshasthiseffect,thatthePhilanthus\'abdomenisinsideandtheBee\'soutside。Thelatter\'sstingthereforefindsunderitspointonlythedorsalsurfaceofthefoe,aconvex,slipperysurfaceandsowellarmouredastobealmostinvulnerable。Thereisherenobreachintowhichtheweaponcanslipbyaccident;andsotheoperationisconductedwithabsolutesurgicalsafety,notwithstandingtheindignantprotestsofthepatient。
Afterthefatalstrokehasbeenadministered,themurderessremainsforalongtimebellytobellywiththedead,forreasonswhichweshallshortlyperceive。TheremaynowbesomedangerforthePhilanthus。Theattitudeofattackanddefenceisabandoned;andtheventralsurface,morevulnerablethantheother,iswithinreachofthesting。Nowthedeceasedstillretainsthereflexuseofherweaponforafewminutes,asIlearnttomycost。HavingtakentheBeetooearlyfromthebanditandhandlingherwithoutsuspectinganyrisk,Ireceivedamostdownrightsting。ThenhowdoesthePhilanthus,inherlongcontactwiththebutcheredBee,managetoprotectherselfagainstthatlancet,whichisbentuponavengingthemurder?Isthereanychanceofacommutationofthedeath-penalty?CananaccidenteverhappenintheBee\'sfavour?Perhaps。
Oneincidentstrengthensmyfaithinthisperhaps。IhadplacedfourBeesandasmanyEristalesunderthebell-glassatthesametime,withtheobjectofestimatingthePhilanthus\'entomologicalknowledgeinthematterofthedistinctionofspecies。Reciprocalquarrelsbreakoutinthemixedcolony。Suddenly,inthemidstofthefray,thekilleriskilled。Shetumblesoveronherback,shewavesherlegs;sheisdead。Whostrucktheblow?ItwascertainlynottheexcitablebutpacificDrone-fly;itwasoneoftheBees,whostruckhomebyaccidentduringthethickofthefight。
Whereandhow?Icannottell。Theincidentoccursonlyonceinmynotes,butitthrowsalightuponthequestion。TheBeeiscapableofwithstandingheradversary;shecanthenandthereslayherwould-beslayerwithathrustofthesting。Thatshedoesnotdefendherselftobetterpurpose,whenshefallsintoherenemy\'sclutches,isduetoherignoranceoffencingandnottotheweaknessofherweapon。Andhereagainarises,moreinsistentlythanbefore,thequestionwhichIaskedabove:howisitthatthePhilanthushaslearntforoffensivewhattheBeehasnotlearntfordefensivepurposes?Iseebutoneanswertothedifficulty:theoneknowswithouthavinglearnt;theotherdoesnotknowbecausesheisincapableoflearning。
LetusnowconsiderthemotivesthatinducethePhilanthustokillherBeeinsteadofparalysingher。Whenthecrimehasbeenperpetrated,shemanipulatesherdeadvictimwithoutlettinggoofitforamoment,holdingitsbellypressedagainstherownsixlegs。Iseeherrecklessly,veryrecklessly,rootingwithhermandiblesinthearticulationoftheneck,sometimesalsointhelargerarticulationofthecorselet,behindthefirstpairoflegs,anarticulationofwhosedelicatemembranesheisperfectlywellaware,eventhough,whenusinghersting,shedidnottakeadvantageofthispoint,whichisthemostreadilyaccessibleofall。Iseeherrough-handlingtheBee\'sbelly,squeezingitagainstherownabdomen,crushingitinthepress。Therecklessnessofthetreatmentisstriking;itshowsthatthereisnoneedforkeepingupprecautions。TheBeeisacorpse;andalittlehustlinghereandtherewillnotdeteriorateitsquality,providedtherebenoeffusionofblood。Inpointoffact,howeverroughthehandling,Ifailtodiscovertheslightestwound。
Thesevariousmanipulations,especiallythesqueezingoftheneck,atoncebringaboutthedesiredresults:thehoneyinthecropmountstotheBee\'sthroat。Iseethetinydropsspurtout,lappedupbythegluttonassoonastheyappear。Thebanditgreedily,overandoveragain,takesthedeadinsect\'slolling,sugaredtongueintohermouth;thensheoncemoredigsintotheneckandthorax,subjectingthehoney-bagtotherenewedpressureofherabdomen。Thesyrupcomesandisinstantlylappedupandlappedupagain。Inthiswaythecontentsofthecropareexhaustedinsmallmouthfuls,yieldedoneatatime。Thisodiousmealattheexpenseofacorpse\'sstomachistakeninasybariticattitude;thePhilanthusliesonhersidewiththeBeebetweenherlegs。Theatrociousbanquetsometimeslastsforhalfanhourorlonger。AtlastthedrainedBeeisdiscarded,notwithoutregret,itseems,forfromtimetotimeIseethemanipulationrenewed。Aftertakingaturnroundthetopofthebell-jar,therobberofthedeadreturnstoherpreyandsqueezesit,lickingitsmouthuntilthelasttraceofhoneyhasdisappeared。
ThisfrenziedpassionofthePhilanthusfortheBee\'ssyrupisdeclaredinyetanotherfashion。Whenthefirstvictimhasbeensuckeddry,Islipundertheglassasecondvictim,whichispromptlystabbedunderthechinandthensubjectedtopressuretoextractthehoney。Athirdfollowsandundergoesthesamefatewithoutsatisfyingthebandit。Iofferafourthandafifth。Theyareallaccepted。MynotesmentiononePhilanthuswhoinfrontofmyeyessacrificedsixBeesinsuccessionandsqueezedouttheircropsintheregulationmanner。Theslaughtercametoanendnotbecausethegluttonwassatedbutbecausemyfunctionsasapurveyorwerebecomingratherdifficult:thedrymonthofAugustcausestheinsectstoavoidmyharmas,whichatthisseasonisdenudedofflowers。Sixcropsemptiedoftheirhoney:whatanorgy!Andeventhentheravenouscreaturewouldverylikelynothavescornedacopiousadditionalcourse,hadIpossessedthemeansofsupplyingit!
Thereisnoreasontoregretthisbreakintheservice;thelittlethatI
havesaidismorethanenoughtoprovethesingularcharacteristicsoftheBee-slayer。IamfarfromdenyingthatthePhilanthushasanhonestmeansofearningherlivelihood;IfindherworkingontheflowersasassiduouslyastheotherWasps,peacefullydrawingherhoneyedbeakers。Themaleseven,possessingnolancet,knownoothermannerofrefreshment。Themothers,withoutneglectingthetabled\'hoteoftheflowers,supportthemselvesbybrigandageaswell。WearetoldoftheSkua,thatpirateoftheseas,thatheswoopsdownuponthefishingbirds,atthemomentwhentheyrisefromthewaterwithacapture。Withablowofthebeakdeliveredinthepitofthestomachhemakesthemgiveuptheirprey,whichiscaughtbytherobberinmid-air。Thedespoiledbirdatleastgetsoffwithnothingworsethanacontusionatthebaseofthethroat。ThePhilanthus,alessscrupulouspirate,pouncesontheBee,stabshertodeathandmakesherdisgorgeinordertofeeduponherhoney。
IsayfeedandIdonotwithdrawtheword。TosupportmystatementIhavebetterreasonsthanthosesetforthabove。InthecagesinwhichvariousHuntingWasps,whosestratagemsofwarIamengagedinstudying,arewaitingtillIhaveprocuredthedesiredprey——notalwaysaneasything——I
haveplantedafewflower-spikes,athistle-headortwo,onwhichareplaceddropsofhoneyrenewedatneed。Heremycaptivescometotaketheirmeals。WiththePhilanthus,theprovisionofhoneyedflowers,thoughfavourablyreceived,isnotindispensable。IhaveonlytoletafewliveBeesintohercagefromtimetotime。Halfadozenadayisabouttheproperallowance。Withnootherfoodthanthesyrupextractedfromtheslain,Ikeepmyinsectsgoingforafortnightorthreeweeks。
Itisasplainasapikestaff:outsidemycages,whentheopportunityoffers,thePhilanthusmustalsokilltheBeeonherownaccount。TheOdynerusasksnothingfromtheChrysomelabutamerecondiment,thearomaticjuiceoftherump;theotherextractsfromhervictimanamplesupplementtohervictuals,thecropfullofhoney。WhatahecatombofBeesmustnotacolonyofthesefreebootersmakefortheirpersonalconsumption,nottomentionthestoredprovisions!IrecommendthePhilanthustothesignalvengeanceofourBee-masters。
Letusgonodeeperintothefirstcausesofthecrime。Letusacceptthingsasweknowthemforthemoment,withtheirapparentorrealatrocity。Tofeedherself,thePhilanthusleviestributeontheBee\'scrop。
Havingmadesureofthis,letusconsiderthebandit\'smethodmoreclosely。
ShedoesnotparalysehercaptureaccordingtotheritescustomaryamongtheHuntingWasps;shekillsit。Whykillit?Iftheeyesofourunderstandingbenotclosed,theneedforsuddendeathisclearasdaylight。ThePhilanthusproposestoobtainthehoneyedbrothwithoutrippinguptheBee,aproceedingwhichwoulddamagethegamewhenitishuntedonbehalfofthelarvae,withoutresortingtothemurderousextirpationofthecrop。Shemust,byablehandling,byskilfulpressure,maketheBeedisgorge,shemustmilkher,inamannerofspeaking。SupposetheBeestungbehindthecorseletandparalysed。Thatdeprivesherofherpoweroflocomotion,butnotofhervitality。Thedigestiveorgansinparticularretainorverynearlyretaintheirnormalenergy,asisprovedbythefrequentexcretionsthattakeplaceintheparalysedprey,solongastheintestineisnotempty,asisprovedaboveallbythevictimsoftheLanguedocianSphex(Cf。“TheHuntingWasp“:chapters8to10——Translator\'sNote。),thosehelplesscreatureswhichIusedtokeepaliveforfortydaysonendwithasoupconsistingofsugarandwater。Itisabsurdtohope,withouttherapeuticmeans,withoutaspecialemetic,tocoaxasoundstomachintoemptyingitscontents。ThestomachoftheBee,whoisjealousofhertreasure,wouldlenditselftotheprocessevenlessreadilythananother。Whenparalysed,theinsectisinert;buttherearealwaysinternalenergiesandorganicforceswhichwillnotyieldtothemanipulator\'spressure。ThePhilanthuswillnibbleatthethroatandsqueezethesidesinvain:thehoneywillnotrisetothemouthsolongasavestigeoflifekeepsthecropclosed。
Thingsaredifferentwithacorpse。Thetensionisrelaxed,themusclesbecomeslack,theresistanceofthestomachceasesandthebagofhoneyisemptiedbytherobber\'svigorouspressure。Yousee,therefore,thatthePhilanthusisexpresslyobligedtoinflictasuddendeath,whichwilldoawayatoncewiththeelasticityoftheorgans。Whereisthelightningstroketobedelivered?Theslayerknowsbetterthanwedo,whenshestickstheBeeunderthechin。Thecerebralgangliaarereachedthroughthelittleholeintheneckanddeathensuesimmediately。
Therelationoftheseactsofbrigandagecannotsatisfymydistressinghabitoffollowingeachreplyobtainedwithafreshquestion,untilthegranitewalloftheunknowablerisesbeforeme。IfthePhilanthusisanexpertinkillingBeesandemptyingcropsswollenwithhoney,thiscannotbemerelyanalimentaryresource,especiallywhen,incommonwiththeothers,shehasthebanqueting-halloftheflowers。Icannotacceptheratrocioustalentasinspiredmerelybythecravingforafeastobtainedattheexpenseofanemptystomach。Somethingcertainlyescapesus:thewhyandwhereforeofthatcropdraineddry。AcreditablemotivemayliehiddenbehindthehorrorswhichIhaverelated。Whatisit?
Anyonecanunderstandthevaguenessoftheobserver\'smindwhenhefirstaskshimselfthisquestion。Thereaderisentitledtobetreatedwithconsideration。Iwillsparehimtherecitalofmysuspicions,mygropingsandmyfailuresandwillcomestraighttotheresultsofmylonginvestigation。Everythinghasitsharmoniousreasonforexistence。IamtoofullypersuadedofthistobelievethatthePhilanthuspursuesherhabitofprofaningcorpsessolelytosatisfyhergreed。Whatdoestheemptiedcropportend?Mayitnotbethat……?Why,yes……Afterall,whoknows?……Letustryalongtheselines。
Themother\'sfirstcareisthewelfareofthefamily。Sofar,wehaveseenthePhilanthushuntingonlyforherstomach\'ssake;letuswatchherhuntingasamother。Nothingiseasierthantodistinguishthetwoperformances。WhentheWaspwantsafewgoodmouthfulsandnothingmore,shescornfullyabandonstheBeeafterpickinghercrop。TheBeeistoheraworthlessremnant,whichwillshrivelwhereitliesandbedissectedbytheAnts。If,ontheotherhand,shewantstostowawaytheBeeasaprovisionforherlarvae,sheclaspsherinhertwointermediatelegsand,walkingontheotherfour,goesroundandroundtheedgeofthebell-glass,seekingforanoutletthroughwhichtoflyoffwithherprey。Whensherecognizesthecirculartrackasimpossible,sheclimbsupthesides,thistimeholdingtheBeebytheantennaewithhermandiblesandclingingtothepolishedandperpendicularsurfacewithhersixfeet。Shereachesthetopoftheglass,staysforalittlewhileinthehollowoftheknobatthetop,returnstotheground,resumeshercirclingandherclimbinganddoesnotdecidetorelinquishherBeeuntilshehasstubbornlyattemptedeverymeansofescape。ThispersistenceonherparttoretainherholdonthecumbrousburdentellsusprettyplainlythatthegamewouldgostraighttothecellsifthePhilanthushadherliberty。
Well,theseBeesintendedforthelarvaearestungunderthechinliketheothers;theyarerealcorpses;theyaremanipulated,squeezed,drainedoftheirhoneyexactlyastheothersare。Inalltheserespects,thereisnodifferencebetweenthehuntconductedtoprovidefoodforthelarvaeandthehuntconductedmerelytogratifythemother\'sappetite。
Astheworriesofcaptivitymightwellbethecauseofafewanomaliesintheinsect\'sactions,IfeltthatIoughttoenquirehowthingshappenintheopen。IlayinwaitnearsomecoloniesofPhilanthi,forlongerperhapsthanthequestiondeserved,asithadalreadybeensettledbywhathadhappenedunderglass。Mytediouswatcheswererewardedfromtimetotime。
Mostofthehuntressesreturnedhomeimmediately,withtheBeeundertheirabdomen;somehaltedonthebrambleshardby;andhereIsawthemsqueezingthedeadBeeandmakingherdisgorgethehoney,whichwasgreedilylappedup。Afterthesepreliminariesthecorpsewasstored。Everydoubtisthereforeremoved:theprovisionsofthelarvaarefirstcarefullydrainedoftheirhoney。
Sinceweareonthespot,letusprolongourstayandenquireintothecustomsofthePhilanthusinastateofliberty。Servingdeadprey,whichgoesbadinafewdays,theBee-huntresscannotadoptthemethodofcertaininsectswhichparalyseanumberofseparateheadsofgameandfillthecellwithprovisions,completingtherationbeforelayingtheegg。SheneedsthemethodoftheBembex,whoselarvareceivesthenecessarynourishmentatintervals,asitgrowslarger。Thefactsconfirmthisdeduction。JustnowI
describedastediousmywatchesnearthecoloniesofthePhilanthi。Theyweretediousinfact,evenmoresoperhapsthanthosewhichtheBembecesusedtoinflictuponmeintheolddays。OutsidetheburrowsoftheGreatCercerisandotherWeevil-lovers,outsidethoseoftheYellow-wingedSphex,theCricket-slayer,thereisplentyofdistraction,thankstothebustlingmovementofthehamlet。Themotherhashardlycomebackhomebeforeshegoesoutagain,soonreturningladenwithanewpreyandoncemoresettingoutuponthechase。Thegoingandcomingisrepeatedatcloseintervalsuntilthewarehouseisfull。
TheburrowofthePhilanthusisfarfromshowinganysuchanimation,eveninapopulouscolony。Invainweremywatchesprolongedforwholemorningsorafternoons;itwasbutveryrarelythatthemotherwhomIhadseengoinwithaBeecameoutagainforasecondexpedition。TwocapturesatmostbythesamehuntresswasallthatIwasabletoseeduringmylongvigils。
Feedingfromdaytodayinvolvesthisdeliberation。Oncethefamilyissuppliedwithasufficientrationforthemoment,themothersuspendsherhunting-tripsuntilfurtherneedarisesandoccupiesherselfwithmining-
workinherundergroundhouse。Cellsaredug;Iseetherubbishgraduallypusheduptothesurface。Beyondthisthereisnotasignofactivity;itisasthoughtheburrowweredeserted。
Theinspectionofthesiteisnoeasymatter。Theshaftdescendstoadepthofnearlythreefeetinacompactsoil,eitherverticallyorhorizontally。
Thespadeandpick,wieldedbystrongerbutlessexperthandsthanmine,areindispensable,forwhichreasontheprocessofexcavationisfarfromsatisfyingmefully。Attheendofthislongtunnel,whichthestrawwhichIuseforsoundingdespairsofeverreaching,thecellsareatlastencountered,ovalcavitieswithahorizontalmajoraxis。Theirnumberandgeneralarrangementescapeme。
Someofthemalreadycontainthecocoon,whichisslenderandsemitransparent,likethoseoftheCerceris,and,likethem,suggeststheshapeofcertainhomoeopathicphials,withovalbelliessurmountedbyataperingneck。Thecocoonisfastenedtotheendofthecellbythetipofthisneck,whichisdarkenedandhardenedbythelarva\'sexcrement;ithasnoothersupport。Itlookslikeashortclubfixedbytheendofthehandlealongthehorizontalaxisofthenest。Othercellscontainthelarvainamoreorlessadvancedstage。Thegrubismunchingthelastmorselservedtoit,withthescrapsofthevictualsalreadyconsumedlyingaroundit。
OtherslastlyshowmeaBee,oneonly,stilluntouchedandbearinganegglaidonherbreast。Thisisthefirstpartialration;theotherswillcomeasandwhenthegrubgrowslarger。Myanticipationsarethusconfirmed:
followingtheexampleoftheBembeces,theFly-killers,thePhilanthus,theBee-killer,layshereggonthefirstpiecewarehousedandatintervalsaddstohernurselings\'repast。
Theproblemofthedeadgameissolved。Thereremainsthisotherproblem,oneofincomparableinterest:whyaretheBeesrobbedoftheirhoneybeforebeingservedtothelarvae?IhavesaidandIsayagainthatthekillingandsqueezingcannotbeexplainedandexcusedsimplybyreferencetothePhilanthus\'loveofgormandizing。Robbingtheworkerofherbootyisnothingoutoftheway:weseeitdaily;butcuttingherthroatinordertoemptyherstomachisgoingbeyondajoke。And,astheBeespackedawayinthecellararesqueezeddryjustasmuchastheothers,thethoughtoccurstomymindthatarumpsteakwithjamisnottoeverybody\'slikingandthatthegamestuffedwithhoneymightwellbeadistastefulorevenunwholesomedishforthePhilanthus\'larvae。Whatwillthegrubdowhen,satedwithbloodandmeat,itfindstheBee\'shoney-bagunderitsmandiblesandespeciallywhen,nibblingatrandom,itripsopenthecropandspoilsitsvenisonwithsyrup?Willitthriveonthemixture?Willthelittleogrepasswithoutrepugnancefromthegamyflavourofacarcasetothescentofflowers?Abluntstatementordenialwouldservenopurpose。Wemustsee。
Letussee。
IrearsomeyoungPhilanthus-grubs,alreadywaxinglarge;but,insteadofsupplyingthemwiththepreytakenfromtheburrows,Igivethemgameofmyowncatching,gamerepletewithnectarfromtherosemaries。MyBees,whomI
killbycrushingtheirheads,arereadilyaccepted;andIatfirstseenothingthatcorrespondswithmysuspicions。Thenmynurselingslanguish,disdaintheirfood,giveacarelessbitehereandthereandendbyperishing,fromthefirsttothelast,besidetheirunfinishedvictuals。
Allmyattemptsmiscarry:Idonotoncesucceedinrearingmylarvaetothestageofspinningthecocoon。AndyetIamnonoviceinthefunctionsofafoster-father。Howmanypupilshavenotpassedthroughmyhandsandreachedmaturityinmyoldsardine-boxesascomfortablyasintheirnaturalburrows!
Iwillnotdrawrashconclusionsfromthischeck;Iamconscientiousenoughtoascribeittoanothercause。Itmaybethattheatmosphereofmystudyandthedrynessofthesandservingasabedhavehadabadeffectonmycharges,whosetenderskinsareaccustomedtothewarmmoistureofthesubsoil。Letusthereforetryanotherexpedient。
ItishardlyfeasibletodecidepositivelybythemethodswhichIhavebeenfollowingwhetherthehoneyisorisnotrepugnanttothegrubsofthePhilanthus。Thefirstmouthfulsconsistofmeat;andthennothingparticularoccurs:itisthenaturaldiet。Thehoneyismetwithlater,whenthemorselhasbeenlargelybitteninto。Ifhesitationandlackofappetitearedisplayedatthisstage,theycometoolateinthedaytobeconclusive:thelarva\'sdiscomfortmaybeduetoother,knownorunknown,causes。Thethingtodowouldbetoofferthegrubhoneyfromthefirst,beforeartificialrearinghasaffecteditsappetite。Itisuseless,ofcourse,tomaketheattemptwithpurehoney:nocarnivorouscreaturewouldtouchit,thoughitwerestarving。Thejam-sandwichistheonlydevicefavourabletomyplans,ameagrejam-sandwich,thatistosay,thedeadBeelightlysmearedorvarnishedwithhoneybymeansofacamel\'s-hairpencil。
Undertheseconditions,theproblemissolvedwiththefirstfewmouthfuls。
Thegrubthathasbittenintothehoneyedpreydrawsbackindisgust,hesitatesalongtimeandthen,urgedbyhunger,beginsagain,triesthissideandthatandendsbyrefusingtotouchthedish。Forafewdaysitpinesawayontopofitsalmostintactprovisions;thenitdies。Allthataresubjectedtothisregimensuccumb。Dotheymerelyperishofinanitioninthepresenceofanunaccustomedfood,whichrevoltstheirappetite,oraretheypoisonedbythesmallquantityofhoneyabsorbedwiththeearlymouthfuls?Icannottell。Thefactremainsthat,whetherpoisonousorrepugnant,theBeeinthestateofbreadandjamisdeathtothem;andthisresultexplains,moreclearlythantheunfavourablecircumstanceofmyformerexperiment,myfailureswiththeBeethathadnotbeenmadetodisgorge。
ThisrefusaltotouchtheunwholesomeordistastefulhoneyisconnectedwithprinciplesofnutritionwhicharetoogeneraltoconstituteagastronomicpeculiarityofthePhilanthus。Theothercarnivorouslarvae,atleastintheorderoftheHymenoptera,areboundtoshareit。Letustry。
Wewillgotoworkasbefore。Iunearththelarvaewhentheyhaveattainedamediumsize,toavoidtheweaknessofinfancy;Itakeawaythenaturalprovisions,smearthecarcasesseparatelywithhoneyand,whenthisisdone,restoreitsvictualstoeachofthegrubs。Ihadtomakeachoice:
noteverysubjectwasequallysuitedtomyexperiments。Imustrejectthelarvaewhicharefedononefatjoint,suchasthoseoftheScolia。Thegrubinfactattacksitspreyatadeterminedpoint,dipsitsheadandneckintotheinsect\'sbody,rootingskilfullyintheentrailstokeepthegamefreshuntiltheendofthemeal,anddoesnotwithdrawfromthebreachuntilthewholeskinisemptiedofitscontents。
Tomakeitletgowiththeobjectofcoatingtheinsideofthevenisonwithhoneyhadtwodrawbacks:Ishouldbecompromisingthelingeringvitalitywhichsavestheinsectthatisbeingdevouredfromgoingbadand,atthesametime,Ishouldbedisturbingthedelicateartofthedevouringinsect,which,ifremovedfromthelodewhichitwasworking,wouldnolongerbeabletorecoveritortodistinguishbetweenthelawfulandtheunlawfulmorsels。ThelarvaoftheScolia,consumingitsCetonia-grub,hastaughtusallthatwewanttoknowonthissubjectinmyearliervolume。(Chapters2
to5ofthepresentvolumecontainthewholeofthematterreferredtoabove——Translator\'sNote。)Theonlyacceptablelarvaearethosesuppliedwithaheapofsmallinsects,whichareattackedwithoutanyspecialart,dismemberedatrandomandeatenupquickly。AmongtheseIhavetestedsuchaschancethrewinmyway:thoseofvariousBembeces,allfedonFlies,thoseofthePalarus,whosebilloffareconsistsofaverylargeassortmentofHymenoptera;thoseoftheTarsalTachytes,suppliedwithyoungLocusts;thoseoftheNest-buildingOdynerus,furnishedwithChrysomela-grubs;thoseoftheSandCerceris,endowedwithapinchofWeevils。Agoodlyvariety,asyousee,ofconsumersandconsumed。Well,toallofthesetheseasoningwithhoneyprovedfatal。Whetherpoisonedordisgusted,theyalldiedinafewdays。
Astrangeresultindeed!Honey,thenectaroftheflowers,thesoledietoftheBee-tribeinbothitsformsandthesoleresourceoftheWaspinheraadultform,istothelarvaeofthelatteranobjectofinsurmountablerepugnanceandprobablyatoxicdish。Eventhetransformationofthenymphosissurprisesmelessthanthisinversionoftheappetite。Whathappensintheinsect\'sstomachtomaketheadultseekpassionatelywhattheyoungsterrefusedlestitshoulddie?Thisisnotaquestionoforganicdebilityunabletoendureatoosubstantial,toohard,toohighlyspiceddish。ThegrubthatgnawstheCetonia-larva,thatgenerouspieceofbutcher\'smeat;thegluttonthatcrunchesitsbatchoftoughLocusts;theonethatbattensonnitrobenzine-flavouredgame:theycertainlyownunfastidiousgulletsandaccommodatingstomachs。Andtheserobusteatersallowthemselvestodieofhungerordigestivetroublesbecauseofadropofsyrup,thelightestfoodimaginable,suitedtotheweaknessofextremeyouthandafeastfortheadultbesides!Whatagulfofobscurityinthestomachofawretchedgrub!
Thesegastronomicalresearchescalledforacounterexperiment。Thecarnivorouslarvaiskilledbyhoney。Conversely,isthemellivorouslarvakilledbyanimalfood?Reservationsareneedfulhere,asintheprevioustests。WeshouldbecourtingaflatrefusalifweofferedapinchofLocuststothelarvaeoftheAnthophoraortheOsmia,forinstance。(ForboththeseWildBeescf。“Bramble-beesandOthers“:passim——Translator\'sNote。)Thehoney-fedinsectwouldnotbiteintoit。Therewouldbenousewhateverintrying。Wemustfindtheequivalentofthejam-sandwichaforesaid;inotherwords,wemustgivethelarvaitsnaturalfarewithamixtureofanimalfood。Theadditionmadebymyartificesshallbealbumen,asfoundintheeggoftheHen,albumentheisomeroffibrin,whichistheessentialfactorinanyformofprey。
Ontheotherhand,theThree-hornedOsmialendsherselfmostadmirablytomyplans,becauseofherdryhoney,consistingforthegreaterpartofflourypollen。Ithereforekneadthishoneywithalbumen,graduatingthedoseuntilitsweightlargelyexceedsthatoftheflour。InthiswayI
obtainpastesofdifferentdegreesofconsistency,butallfirmenoughtobearthelarvawithoutdangerofimmersion。Withtoofluidamixturetherewouldbeariskofdeathbydrowning。LastlyIinstallamoderately-
developedlarvaoneachofmyalbuminouscakes。
Thedishofmyinventingdoesnotincitedislike:farfromit。Thegrubsattackitwithouthesitationandconsumeitwitheveryappearanceoftheusualappetite。Thingscouldnotgobetterifthefoodhadnotbeenalteredbymyculinaryrecipes。Everythinggoesdown,includingthemorselsinwhichIfearedthatIhadoverdonetheadditionofalbumen。And——anevenmoreimportantpoint——theOsmia-larvaefedinthismannerattaintheirnormaldimensionsandspintheircocoons,fromwhichadultinsectsissueinthefollowingyear。Notwithstandingthealbuminousregimen,thecycleoftheevolutionisachievedwithoutimpediment。
Whatarewetoconcludefromallthis?Ifeelgreatlyembarrassed。Omnevivumexovo,thephysiologiststellus。Everyanimaliscarnivorous,initsfirstbeginnings:itisformedandnourishedatthecostofitsegg,inwhichalbumenpredominates。Thehighest,themammal,adherestothisdietforalongtime:ithasitsmother\'smilk,richincasein,anotherisomerofalbumen。Thegramnivorousnestlingisfirstfedongrubs,whicharebetteradaptedtothenicetiesofitsstomach;manyoftheminutestnew-
borncreatures,beingatoncelefttotheirowndevices,taketoanimalfood。Inthiswaytheoriginalmethodofnourishmentiscontinuedforallalike:themethodwhichallowsfleshtobemadefromfleshandbloodfromblood,withnochemicalprocessbeyondthesimplestmodification。Atmaturity,whenthestomachhasacquireditsfullstrength,vegetablefoodisadopted,involvingamorecomplicatedchemistrybuteasiertoobtain。
Milkisfollowedbyfodder,wormsbyseeds,thepreyintheburrowbythenectaroftheflowers。
ThissuppliesapartialexplanationofthetwofolddietoftheHymenopterawithcarnivorouslarvae:meatfirst,honeynext。Butthenthenoteofinterrogationisshifted。Itstoodelsewhere;itnowstandshere。WhyistheOsmia,whoasalarvafaressowellonalbumen,fedonhoneyatthestart?WhydotheBee-tribereceiveavegetabledietwhentheothermembersoftheorderreceiveananimaldiet?
IfIwereabelieverinevolution,Ishouldsayyes,bythefactofitsgerm,everyanimalisoriginallycarnivorous。Theinsectinparticularstartswithalbuminoidmaterials。Manylarvaeadheretotheegg-food,manyadultinsectsdolikewise。Butthestruggletofillthebelly,whichafterallisthestruggleforlife,demandssomethingbetterthantheprecarioushazardsofthechase。Man,atfirstaravenoushunteraftergame,broughttheflockintoexistenceandturnedshepherdtoavoidatimeofdearth。Anevengreaterprogressinspiredhimtoscrapetheearthandtosowseed,whichassureshimofaliving。Theevolutionfromscarcitytomoderationandfrommoderationtoplentyhasledtotheresourcesofhusbandry。
Theanimalsforestalledusthispathofprogress。TheancestorsofthePhilanthus,intheremoteagesofthelacustriantertiaryformations,livedbypreyinboththelarvalandtheadultforms:theyhuntedforthemselvesaswellasforthefamily。TheydidnotconfinethemselvestoemptyingtheBee\'scrop,astheirdescendantsdotothisday:theydevouredthedeceased。Fromthebeginningtotheendtheyremainedflesh-eaters。Later,fortunateinnovators,whoseracesupplantedthelaggards,discoveredaninexhaustiblenourishment,obtainedwithoutdangerousconflictsorlaborioussearch:thesugarysecretionsoftheflowers。Thecostlyhabitoflivingonprey,whichdoesnotfavourlargepopulations,wasmaintainedforthefeeblelarvae;butthevigorousadultbrokeherselfofittoleadaneasierandmoreprosperouslife。Thus,gradually,wasformedthePhilanthusofourday;thuswasacquiredthetwofolddietofthevariouspredatoryinsectsourcontemporaries。
TheBeehasdonebetterstill:fromthemomentofleavingtheeggshedeliveredherselfcompletelyfromfood-stuffstheacquisitionofwhichdependedonchance。Shediscoveredhoney,thegrubs\'food。Renouncingthechaseforeverandbecominganagriculturalistpureandsimple,theinsectattainsadegreeofphysicalandmoralprosperitywhichthepredatoryspeciesarefarfromsharing。HencetheflourishingcoloniesoftheAnthophorae,theOsmiae,theEucerae(Agenusoflong-hornedBurrowingBees——Translator\'sNote。),theHalictiandotherhoney-manufacturers,whereasthepredatoryinsectsworkinisolation;hencethesocietiesinwhichtheBeedisplaysherwonderfultendencies,thesupremeexpressionofinstinct。
ThisiswhatIshouldsayifIbelongedtothatschool。Itallformsachainofverylogicaldeductionsandproffersitselfwithacertainairoflikelihoodwhichweshouldbegladtofindinahostofevolutionistargumentsputforwardasirrefutable。Well,Iwillmakeapresentofmydeductiveviews,withoutregret,towhoevercarestohavethem:Idon\'tbelieveonewordofthem;andIconfessmyprofoundignoranceoftheoriginofthetwofolddiet。
WhatIdounderstandmoreclearly,afteralltheseinvestigations,isthetacticsofthePhilanthus。Whenwitnessingherferociousfeasting,therealreasonofwhichwasunknowntome,Iheapedthemostill-soundingepithetsuponher,callingheramurderess,abandit,apirate,arobberofthedead。Ignoranceisalwaysevil-tongued;themanwhodoesnotknowindulgesinrudeassertionsandmischievousinterpretations。Nowthatmyeyeshavebeenopenedtothefacts,IhastentoapologizeandtorestorethePhilanthustoherplaceinmyesteem。IndrainingthecropsofherBeesthemotherisperformingthemostpraiseworthyofallactions:sheisprotectingherfamilyagainstpoison。Ifshehappenstokillonherownaccountandtoabandonthecorpseaftermakingitdisgorge,Idarenotreckonthisagainstherasacrime。WhenthehabithasbeenformedofemptyingtheBee\'scropwithagoodmotive,thereisagreattemptationtodoitagainwithnootherexcusethanhunger。Besides,whoknows?Perhapsthereisalwaysatthebackofherhuntingsomethoughtofgamewhichmightbeusefulforthelarvae。Althoughnotcarriedintoeffect,theintentionexcusesthedeed。
Ithereforewithdrawmyepithetsinordertoadmiretheinsect\'smaternallogicandtoholdituptotheadmirationofothers。Thehoneywouldbepernicioustothehealthofthelarvae。Howdoesthemotherknowthatthesyrup,atreatforher,isunwholesomeforheryoung?Tothisquestionourscienceoffersnoreply。Thehoney,Isay,wouldimperilthegrubs\'lives,TheBeemustthereforefirstbemadetodisgorge。Thedisgorgingmustbeeffectedwithoutlaceratingthevictim,whichthenurselingmustreceiveinthefreshstate;andtheoperationisimpracticableonaparalysedinsectbecauseoftheresistanceofthestomach。TheBeemustthereforebekilledoutrightinsteadofbeingparalysed,orthehoneywillnotbevoided。
Instantaneousdeathcanbeinflictedonlybywoundingtheprimordialcentreoflife。Thestingmustthereforeaimatthecervicalganglia,theseatofinnervationonwhichtherestoftheorganismdepends。Toreachthemthereisonlyoneway,throughthelittlegapinthethroat。Itisherethereforethatthestingmustbeinserted;anditishereinfactthatitisinserted,inaspothardlyaslargeasthetwenty-fifthofaninchsquare。
Suppressasinglelinkofthiscompactchain,andtheBee-fedPhilanthusbecomesimpossible。
Thathoneyisfataltocarnivorouslarvaeisafactwhichteemswithconsequences。SeveralHuntingWaspsfeedtheirfamiliesuponBees。Theseinclude,tomyknowledge,theCrownedPhilanthus(P。coronatus,FAB。),wholinesherburrowswithbigHalicti;theRobberPhilanthus(P。raptor,LEP。),whochasesallthesmaller-sizedHalicti,suitedtoherowndimensions,indifferently;theOrnateCerceris(C。ornata,FAB。),anotherpassionateloverofHalicti;andthePalarus(P。flavipes,FAB。),who,withacuriouseclecticism,stacksinhercellsthegreaterpartoftheHymenopteronclanthatdoesnotexceedherpowers。Whatdothesefourhuntressesandtheothersofsimilarhabitsdowiththeirvictimswhosecropsaremoreorlessswollenwithhoney?TheymustfollowtheexampleoftheBee-eatingPhilanthusandmakethemdisgorge,lesttheirfamilyperishofahoneyeddiet;theymustmanipulatethedeadBee,squeezeheranddrainherdry。Everythinggoestoshowit。Ileaveittothefuturetodisplaythesedazzlingproofsofmydoctrineintheirproperlight。
CHAPTER11。THEMETHODOFTHEAMMOPHILAE。(FortheseSand-wasps,cf。“TheHuntingWasps“:chapters13and18to20——Translator\'sNote。)
Myreadersmaydifferinappraisingthecomparativevalueofthetriflingdiscoverieswhichentomologyowestomylabours。Thegeologist,therecorderofforms,willpreferthehypermetamorphosisoftheOil-beetles(ThechaptertreatingofthissubjecthasnotyetbeentranslatedintoEnglishandwillappearinalatervolume——Translator\'sNote。),thedevelopmentoftheAnthrax(Cf。“TheLifeoftheFly“:chapter2——
Translator\'sNote。)orlarvaldimorphism;theembryogenist,searchingintothemysteriesoftheegg,willhavesomeesteemformyenquiriesintotheegg-layinghabitsoftheOsmia(Cf。“Bramble-beesandOthers“:chapter4——
Translator\'sNote。);thephilosopher,rackinghisbrainoverthenatureofinstinct,willawardthepalmtotheoperationsoftheHuntingWasps。I
agreewiththephilosopher。Withouthesitation,Iwouldabandonalltherestofmyentomologicalbaggageforthisdiscovery,whichhappenstobetheearliestindateandthatofwhichIhavethefondestmemories。NowheredoIfindamorebrilliant,morelucid,moreeloquentproofoftheintuitivewisdomofinstinct;nowheredoesthetheoryofevolutionsufferamoreobstinatecheck。
Darwin,atruejudge,madenomistakeaboutit。(CharlesRobertDarwin,bornthe12thofFebruary,1809,atShrewsbury,diedatDown,inKent,onthe19thofApril,1882。Foranaccountofcertainexperimentswhichtheauthorconductedonhisbehalf,cf。“TheMason-bees“:chapter4——
Translator\'sNote。)Hegreatlydreadedtheproblemoftheinstincts。Myfirstresultsinparticularlefthimveryanxious。IfhehadknownthetacticsoftheHairyAmmophila,theMantis-huntingTachytes,theBee-eatingPhilanthus,theCalicurgiandothermarauders,hisanxiety,Ibelieve,wouldhaveendedinafrankadmissionthathewasunabletosqueezeinstinctintothemouldofhisformula。Alas,thephilosopherofDownquittedthisworldwhenthediscussion,withexperimentstosupportit,hadbarelybegun:amethodsuperiortoanyargument!ThelittlethatIhadpublishedatthattimelefthimwithstillsomehopeofanexplanation。Inhiseyes,instinctwasalwaysanacquiredhabit。ThepredatoryWaspskilledtheirpreyatfirstbystabbingitatrandom,hereandthere,inthesoftestparts。Bydegreestheyfoundthespotwherethestingwasmosteffectual;andthehabitonceformedbecameatrueinstinct。Transitionsfromonemethodofoperationtotheother,intermediarychanges,sufficedtobolsterupthesesweepingassertions。Inaletterofthe16thofApril,1881,heasksG。J。Romanestoconsidertheproblem:
“Idonotknow,“hesays“whetheryouwilldiscussinyourbookonthemindofanimalsanyofthemorecomplexandwonderfulinstincts。Itisunsatisfactorywork,astherecanbenofossilisedinstincts,andthesoleguideistheirstateinothermembersofthesameorder,andmerePROBABILITY。
“Butifyoudodiscussany(anditwillperhapsbeexpectedofyou),I
shouldthinkthatyoucouldnotselectabettercasethanthatofthesand-
waspswhichparalysetheirpreyasdescribedbyFabreinhiswonderfulpaperinthe“Analesdessciencesnaturelles,“andsinceamplifiedinhisadmirable“Souvenirs……“
Ithankyou,Oillustriousmaster,foryoureulogisticexpressions,provingthekeeninterestwhichyoutookinmystudiesofinstinct,noungratefultask——farfromit——whenwetackleitasitshouldbetackled:fromthefront,withtheaidoffacts,andnotfromtheflank,withtheaidofarguments。Argumentsarehereoutofplace,ifwewishtomaintainourpositioninthelight。Besides,wherewouldtheyleadus?Toevokingtheinstinctsofbygoneages,whichhavenotbeenpreservedbyfossilization?
Anysuchappealtothedimanddistantpastisquiteunnecessary,ifwewishforvariationsofinstinct,leadingbydegrees,accordingtoyou,fromoneinstincttoanother;thepresentworldoffersusplenty。
Eachoperatorhasherparticularmethod,herparticularkindofgame,herparticularpointsofattackandtricksoffence;butinthemidstofthisvarietyoftalentsweobserve,immutableandpredominant,theperfectaccordanceofthesurgerywiththevictim\'sorganizationandthelarva\'sneeds。Theartofonewillnotexplaintheartofanother,nolessexactinthedelicacyofitsrules。Eachoperatorhasherowntactics,whichtoleratenoapprenticeship。TheAmmophila,theScolia,thePhilanthusandtheothersalltellusthesamething:nonecanleavedescendantsifshebenotfromtheoutsettheskilfulparalyserorslayerthatsheisto-day。The“almost“isimpracticablewhenthefutureoftheraceisatstake。Whatwouldhavebecomeofthefirst-bornmammalbutforitsperfectinstinctofsuckling?
Andthen,tosupposetheimpossible:aWaspdiscoversbychancetheoperativemethodwhichwillbethesavingattributeofherrace。Howarewetoadmitthatthisfortuitousact,towhichthemotherhasvouchsafednomoreattentionthantoherotherlessfortunateattempts,couldleaveaprofoundtracebehinditandbefaithfullytransmittedbyheredity?Isitnotgoingbeyondreason,goingbeyondthelittlethatisknowntousascertain,ifwegranttoatavismthisstrangepower,ofwhichourpresentworldknowsnoinstance?Thereisagooddealtobesaidforthispointofview,myreveredmaster!But,oncemore,argumentsarehereoutofplace;
thereisroomonlyforfacts,ofwhichIwillresumetherecital。
HithertoIhadbutonemeansofstudyingtheoperativemethodsofthespoilers:tosurprisetheWaspinpossessionofhercapture,torobherofherpreyandimmediatelytogiveherinexchangeasimilarprey,butalivingone。Thismethodofsubstitutionisanexcellentexpedient。Itsonlydefect——averygraveone——isthatitsubjectsobservationtoveryuncertainchances。Thereislittleprospectofmeetingtheinsectdraggingitsvictimalong;and,inthesecondplace,shouldgoodfortunesuddenlysmileuponyou,preoccupiedasyouarewithothermattersyouhavenotthesubstituteathand。Ifweprovideourselveswiththenecessaryheadofgameinadvance,thehuntressisnotthere。Weavoidonereeftofounderonanother。Moreover,theseunlookedforobservations,madesometimesonthepublichighway,theworstoflaboratories,areonlyhalf-satisfactory。Inthecaseofswiftly-enactedscenes,whichitisnotinourpowertorenewagainandagainuntilperfectconvictionisreached,wealwaysfearlestwemaynothaveseenaccurately,maynothaveseeneverything。
Amethodwhichcouldbecontrolledatwillwouldofferthebestguarantees,aboveallifemployedathome,undercomfortableconditions,favourabletoprecision。Iwished,therefore,toseemyinsectsatworkontheactualtableatwhichIamwritingtheirhistory。Hereveryfewoftheirsecretswouldescapeme。Thiswishofminewasanoldone。Asabeginner,ImadesomeexperimentsunderglasswiththeGreatCerceris(C。tuberculata)andtheYellow-wingedSphex。Neitherofthemrespondedtomydesires。TherefusalofeachtoattackrespectivelyherCleonusorherCricketdiscouragedfurtherprogressinthisdirection。Iwaswrongtoabandonmyattemptssosoon。Now,verylongafterwards,theideaoccurstometoplaceunderglasstheBee-eatingPhilanthus,whomIsometimessurpriseintheopenengagedinforcingabeetodisgorgeherhoney。Thecaptivemassacresherbeesinsuchaspiritedfashionthattheoldhoperevivesstrongerthanever。Icontemplatereviewingallthewieldersofthestilettoandforcingeachtorevealhertactics。
Iwasobligedtoabatetheseambitionsconsiderably。Ihadsomesuccessesandmanymorefailures。Iwilltellyouoftheformer。Myinsect-cageisaspaciousdomeofwire-gauzerestingonabedofsand。HereIkeepinreservethecaptivesofmyhunting-expeditions。Ifeedthemonhoney,placedinlittledropsonspikesoflavender,onheadsofthistle,orfielderyngo,orglobe-thistle,accordingtotheseason。Mostofmyprisonersdowellonthisdietandseemscarcelyaffectedbytheirinternment;otherspineawayanddieintwoorthreedays。Thesevictimsofdespairnearlyalwaysthrowmeback,becauseofthedifficultyofobtainingthenecessarypreyatshortnotice。
Indeeditentailsnosmalltroubletosecureinthenickoftimethegamedemandedbythehuntresswhohasrecentlyfallenacaptivetomynet。Asassistant-purveyorsIhaveafewsmallschoolboys,who,releasedfromthetediumoftheirdeclensionsandconjugations,setout,onleavingtheclassroom,toinspectthegreenswardsandbeatthebushesintheneighbourhoodonmybehalf。Thegrossou,thepenny-piece,ifyouplease,stimulatestheirzeal;butwithmisadventurousresults!WhatIneedto-dayisCrickets。ThebandsalliesforthandreturnswithnotasingleCricket,butnumbersofEphippigers,forwhichIaskedthedaybeforeyesterdayandwhichInolongerneed,myLanguedocianSphexbeingdead。Generalsurpriseatthissuddenchangeofmarket。Myyoungscatterbrainsfindithardtounderstandthatthebeastwhichwassoprecioustwodaysagoisnowofnovaluewhatever。When,owingtothechancesofmynet,areneweddemandfortheEphippigersetsin,thentheywillbringmetheCricket,thedespisedCricket。
Suchatradecouldneverholdoutifnowandagainmyspeculatorswerenotencouragedbysomesuccess。Atthemomentwhenurgentnecessityissendingupprices,oneofthembringsmeamagnificentGad-flyintendedfortheBembex。Fortwohours,whenthesunwasatitsheight,hekeptwatchonthethreshing-floorhardby,waitingfortheblood-sucker,inordertocatchhimonthebuttocksoftheMuleswhichtrotroundandroundtramplingthecorn。Thisgallantfellowshallhavehisgrossouandasliceofbreadandjamaswell。Asecond,nolessfortunate,hasfoundafatSpider,theEpeira,forwhommyPompiliarewaiting。TothetwosousofthisfortunateyouthIaddalittlepictureforhismissal。Thusaremypurveyorskeptgoing;and,afterall,theirhelpwouldbeveryinadequateifIdidnottakeuponmyselfthemainburdenofthesewearisomequests。
Onceinpossessionoftherequisiteprey,Itransferthehuntressfrommywarehouse,thewire-gauzecage,toabell-glassvaryingincapacityfromonetothreeorfourlitres(13/4to5or7pints——Translator\'sNote。),accordingtothesizeandhabitsofthecombatants;Iplacethevictiminthearena;Iexposethebell-glasstothedirectraysofthesun,withoutwhichconditiontheexecutionerasaruledeclinestooperate;Iarmmyselfwithpatienceandawaitevents。
WewillbeginwiththeHairyAmmophila,myneighbour。Yearafteryear,whenAprilcomes,Iseeherinconsiderablenumbers,verybusyonthepathsinmyenclosure。UntilJuneIseeherdiggingherburrowsandsearchingfortheGreyWorm,tobeplacedinthemeat-cellar。HertacticsarethemostcomplexthatIknowandmorethananyotherdeservestobethoroughlystudied。Tocapturethecunningvivisector,toreleaseherandcatchheragainIfindaneasymatterforthebestpartofamonth;sheworksoutsidemydoor。
IhavestilltoobtaintheGreyWorm。ThismeansarepetitionofthedisappointmentswhichIhadbefore,when,tofindacaterpillar,IwasobligedtowatchtheAmmophilawhilehuntingandtobeguidedbyherhints,asthetruffle-hunterisguidedbythescentofhisDog。Apatientexplorationoftheharmas,onetuftofthymeafteranother,doesnotgivemeasingleworm。Myrivalsinthissearcharefindingtheirgameateverymoment;Icannotfinditevenonce。Yetonemorereasonforbowingtothesuperiorityoftheinsectinthemanagementofheraffairs。Mybandofschoolboysgettoworkinthesurroundingfields。Nothing,alwaysnothing!
Iinmyturnexploretheouterworld;andfortendaysthepursuitofacaterpillartormentsmetillIlosemypowerofsleep。Then,atlast,victory!Atthefootofasunnywall,underthebuddingrosettesofthepanicledcentaury,IfindafairsupplyofthepreciousGreyWormoritsequivalent。
BeholdthewormandtheAmmophilafacetofacebeneaththebell-glass。
Usuallytheattackispromptenough。Thecaterpillarisgrabbedbytheneckwiththemandibles,wide,curvedpincerscapableofembracingthegreaterpartofthelivingcylinder。Thecreaturethusseizedtwistsandturnsandsometimes,withablowofitstail,sendstheassailantrollingtoadistance。Thelatterisunconcernedandthrustsherstingthriceinrapidsuccessionintothethorax,beginningwiththethirdsegmentandendingwiththefirst,wheretheweaponisdrivenhomewithgreaterdeterminationthanelsewhere。
Thecaterpillaristhenreleased。TheAmmophilastampsontheground;withherquiveringtarsishetapsthecardboardonwhichthebell-glassstands;
sheliesdownflat,dragsherselfalong,getsupagain,flattensherselfoncemore。Thewingsjerkconvulsively。Fromtimetotimetheinsectplacesitsmandiblesandforeheadontheground,thenrearshighuponitshind-
legsasthoughtoturnheadoverheels。InallthisIseeamanifestationofdelight。Werubourhandswhenrejoicingatasuccess;theAmmophilaiscelebratinghertriumphoverthemonsterinherownfashion。Duringthisfitofdeliriousjoy,whatisthewoundedcaterpillardoing?Itcannolongerwalk;butallthepartbehindthethoraxstrugglesviolently,curlinganduncurlingwhentheAmmophilasetsafootuponit。Themandiblesopenandshutmenacingly。
SECONDACT——Whentheoperationisresumed,thecaterpillarisseizedbytheback。Fromfronttorear,inorder,allthesegmentsarestungontheventralsurface,exceptthethreeoperatedon。Allseriousdangerisavertedbythestabsofthefirstact;therefore,theWaspisnowabletoworkuponherpatientwithoutthehastedisplayedattheoutset。
Deliberatelyandmethodicallyshedrivesinherlancet,withdrawsit,selectsthespot,stabsitandbeginsagain,passingfromsegmenttosegment,takingcare,eachtime,tolayholdofthebackalittlemoretotherear,inordertobringthesegmenttobeparalysedwithinreachoftheneedle。Forthesecondtime,thecaterpillarisreleased。Itisabsolutelyinert,exceptthemandibles,whicharestillcapableofbiting。
THIRDACT——TheAmmophilaclaspstheparalysedvictimbetweenherlegs;
withthehooksofhermandiblessheseizesthebackofitsneck,atthebaseofthefirstthoracicsegment。Fornearlytenminutesshemunchesthisweakspot,whichliesclosetothecerebralnerve-centres。Thepincerssqueezesuddenlybutatintervalsandmethodically,asthoughthemanipulatorwishedeachtimetojudgeoftheeffectproduced;thesqueezesarerepeateduntilIamtiredoftryingtocountthem。Whentheycease,thecaterpillar\'smandiblesaremotionless。Thencomesthetransportationofthecarcase,adetailwhichisnotrelevantinthisplace。
Ihavesetforththecompletetragedy,asitisfairlyoftenenacted,butnotalways。Theinsectisnotamachine,unvaryingintheeffectofitsmechanism;itisallowedacertainlatitude,enablingittocopewiththeeventualitiesofthemoment。AnyoneexpectingtoseetheincidentsofthestruggleunfoldingthemselvesexactlyasIhavedescribedwillriskdisappointment。Specialinstancesoccur——theyareevennumerous——whicharemoreorlessatvariancewiththegeneralrule。Itwillbewelltomentionthemoreimportant,inordertoputfutureobserversontheirguard。
Notinfrequentlythefirstact,thatofparalysingthethorax,isrestrictedtotwothrustsofthestinginsteadofthree,oreventoone,whichisthendeliveredintheforemostsegment。This,itwouldseem,fromthepersistencywithwhichtheAmmophilainflictsit,isthemostimportantprickofall。Isitunreasonabletosupposethattheoperator,whenshebeginsbyprickingthethorax,intendstosubduehercaptureandtomakeitincapableofinjuringher,orevenofdisturbingherwhenthemomentcomesforthedelicateandprotractedsurgeryofthesecondact?Thisideaseemstomehighlyadmissible;andthen,insteadofthreedagger-thrusts,whynottwoonly,whynotmerelyone,ifthiswouldsufficeforthetimebeing?Theamountofvigourdisplayedbythecaterpillarmustbetakenintoconsideration。Bethisasitmay,thesegmentssparedinthefirstactarestabbedinthesecond。Ihavesometimesevenseenthethreethoracicsegmentsstungtwiceover:atthebeginningoftheattackandagainwhentheWaspreturnedtohervanquishedprey。
TheAmmophila\'striumphanttransportsbesideherwoundedandwrithingvictimarealsosubjecttoexceptions。Sometimes,withoutreleasingitspreyforamoment,theinsectproceedsfromthethoraxtothenextsegmentsandcompletesitsoperationinasinglespell。Thejoyousentr\'actedoesnottakeplace;theconvulsivemovementsofthewingsandtheacrobaticposturesaresuppressed。
Theruleisparalysisofallthesegments,howevermany,inregularorderfromfronttoback,includingeventheanalsegmentifthisboastoflegs。
Byafairlyfrequentexceptionthelasttwoorthreesegmentsarespared。
Anotherexception,butaveryrareone,ofwhichIhaveobservedonlyasingleinstance,consistsintheinversionofthedagger-thrustsofthesecondact,thethrustsbeingdeliveredfrombacktofront。Thecaterpillaristhenseizedbyitshinderextremity;andtheAmmophila,progressingtowardsthehead,stingsinreverseorder,passingfromthesucceedingtotheprecedingsegment,includingthethoraxalreadystabbed。ThisreversaloftheusualtacticsIaminclinedtoattributetonegligenceontheinsect\'spart。Negligenceornot,theinvertedmethodhasthesamefinalresultasthedirectmethod:theparalysisofallthesegments。
Lastly,thecompressionoftheneckbythemandibularypincers,themunchingoftheweakspotbetweenthebaseoftheskullandthefirstsegmentofthethorax,issometimespractisedandsometimesneglected。Ifthecaterpillar\'sjawsopenandthreaten,theAmmophilastillsthembybitingtheneck;iftheyarealreadygrowingquiescent,sherefrains。
Withoutbeingindispensable,thisoperationisusefulatthemomentofcartingtheprey。Thecaterpillar,tooheavytobecarriedonthewing,isdragged,headfirst,betweentheAmmophila\'slegs。Ifthemandiblesareworking,theleastclumsinessmayrenderthemdangeroustothecarrier,whoisexposedtotheirbitewithoutanymeansofdefence。
Moreover,onceontheway,thicketsofgrassaretraversedinwhichtheGreyWormcanseizeabladeandofferadesperateresistancetothetraction。Noristhisall。TheAmmophiladoesnotasaruletroubleaboutherburrow,oratleastdoesnotcompleteit,untilshehascaughthercaterpillar。Duringthemining-operations,thegameislaidsomewherehighup,outofreachoftheAnts,onsometuftofgrass,orthetwigsofashrub,whitherthehuntress,fromtimetotime,stoppingherwell-sinking,hastenstoseeifherquarryisstillthere。Forherthisisameansofrefreshinghermemoryofthespotwhereshehaslaidit,oftenatsomedistancefromtheburrow,andofpreventingattemptsatrobbery。Whenthemomentcomesforremovingthegamefromitshiding-place,thedifficultywouldbeinsurmountableweretheworm,grippingtheshrubwithallthemightofitsjaws,toanchoritselfthere。Henceinertiaofthepowerfulhooks,whicharetheparalysedcreature\'ssolemeansofresistance,becomesessentialduringthecarting。TheAmmophilaobtainsitbycompressingthecerebralganglia,bymunchingtheneck。Theinertiaistemporary;itwearsoffsoonerorlater;butbythistimethecarcaseisinthecellandtheegg,prudentlylaidatadistanceontheventralsurfaceoftheworm,hasnothingtofearfromthecaterpillar\'sgrapnels。NocomparisonispermissiblebetweenthemethodicalsqueezesoftheAmmophilabenumbingthecephalicnerve-centresandthebrutalmanipulationsofthePhilanthusemptyingthecropofherBee。ThehuntressofGreyWormsinducesatemporarytorporofthemandibles;theravisherofBeesmakesthemejecttheirhoney。Noonegiftedwiththeleastperspicacitywillconfoundthetwooperations。
ForthemomentwewillnotdwellanylongeronthemethodoftheHairyAmmophila;wewillseeinsteadhowherkinswomenbehave。AfterprotractedrefusalstheSandyAmmophila(A。sabulosa,FAB。),onwhomIexperimentedinSeptember,endedbyacceptingtheprofferedprey,apowerfulcaterpillarasthickasalead-pencil。ThesurgicalmethoddidnotdifferfromthatemployedbytheHairyAmmophilawhenoperatingonherGreyWorminonespell。Allthesegments,exceptingthelastthree,werestungfromfronttoback,beginningwiththeprothorax。Thissinglesuccesswithasimplifiedmethodleftmeinignoranceoftheaccessorymanoeuvres,whichIdonotdoubtmustmoreorlesscloselyrecallthoseoftheprecedingspecies。
Iamallthemoreinclinedtoacceptthesesecondarymanoeuvres,notasyetrecorded——thetransportsoftriumphandthecompressionsoftheneck——
inasmuchasIseethempractisedupontheLoopercaterpillars,whichdiffersogreatlyfromtheothersinexternalstructure,exactlyasIhavedescribedtheminthecaseoftheGreyWorm,whichisoftheordinaryform。
Twospecies,theSilkyAmmophila(A。holoserica,FAB。)andJules\'Ammophila(Seeinthefirstvolumeofthe“Souvenirsentomologiques“whatImeanbythisdenomination——Author\'sNote。),affectthiscuriousprey,whichmoveswiththestrideofapairofcompasses。Thefirst,oftenrenewedunderglassduringthegreaterpartofAugust,hasalwaysrefusedmyoffers;thesecond,hercontemporary,has,onthecontrary,promptlyacceptedthem。
IpresentJules\'Ammophilawithaslender,brownishLooperwhichIcaughtonthejasmine。Theattackisnotslowincoming。Thecaterpillarisgrabbedbytheneck:livelycontortionsofthevictim,whichrollstheaggressoroveranddragsheralong,nowuppermost,nowundermostinthestruggle。Firstthethoraxisstung,initsthreerings,frombacktofront。Thestinglingerslongestnearthethroat,inthefirstsegment。
Thisdone,theAmmophilareleaseshervictimandproceedstostamphertarsi,topolishherwings,tostretchherself。AgainIobservetheacrobaticpostures,theforeheadtouchingtheground,thehinderpartofthebodyraised。ThismimictriumphisthesameasthatofthehuntressoftheGreyWorm。ThentheLooperisoncemoreseized。Despiteitscontortions,whicharenotintheleastabatedbythethreewoundsinthethorax,itisstungfromfronttobackineachsegmentstillunwounded,nomatterhowmany,whethersuppliedwithlegsornot。Iexpectedtoseethestingrefrainmoreorlessinthelongintervalwhichseparatesthetruelegsinfrontfromthepro-legsattheback(Fleshylegsfoundontheabdominalsegmentsofcaterpillarsandcertainotherlarvae——Translator\'sNote。):segmentsdevoidoforgansofdefenceorlocomotiondidnotseemtometodeserveconscientioussurgery。Iwasmistaken:notasegmentoftheLooperisspared,noteventhelastones。Itistruethatthese,beingeminentlycapableofcatchingholdwiththeirfalselegs,wouldbedangerouslaterweretheWasptoneglectthem。
Iobserve,however,thatthelancetworksmorerapidlyinthesecondpartoftheoperationthaninthefirst,eitherbecausethecaterpillar,halfsubjugatedbythetriplewoundattheoutset,iseasiertoreachwiththesting,orbecausethesegmentsmoreremotefromtheheadarerenderedharmlesswithasmallerinjectionofpoison。Nowheredoweseerepeatedthecareexpendeduponparalysingthethorax,stilllesstheinsistentattentiontothefirstsegment。OnreturningtoherLooperaftertheentr\'actedevotedtothejoysofsuccess,theAmmophilastabssoswiftlythat,ononeoccasion,Isawherobligedtobeginalloveragain。Lightlystungalongitswholelength,thevictimstillstruggles。Withouthesitation,theoperatorunsheathesherscalpelforthesecondtimeandoperatesontheLooperafresh,withtheexceptionofthethorax,whichwasalreadysufficientlyanaesthetized。Thisdone,allisinorder;thereisnomoremovement。
Afterthestilettothehooksofthemandiblesrarelyfailtointervene。
Longandcurved,theynibbleattheparalysedvictim\'sneck,sometimesfromabove,sometimesfrombelow。ItisarepetitionofwhattheHairyAmmophilashowedus:thesamesuddensqueezesofthepincers,withratherlongintervalsbetween。Theseintervals,thesemeasuredbitesandtheinsect\'swatchfulattitudehaveeveryappearanceoftellingusthattheoperatorisnotingtheeffectproducedbeforegivingafreshpinchofthenippers。
ItwillbeseenhowvaluableistheevidenceofJules\'Ammophila:ittellsusthattheimmolatersofLoopercaterpillarsandthoseofordinarycaterpillarsfollowpreciselythesamemethod;thatvictimsdisplayingverydissimilarexternalstructuredonotentailanymodificationoftheoperativetacticssolongastheinternalorganizationremainsthesame。
Thenumber,arrangementanddegreeofmutualindependenceofthenerve-
centresguidethesting;theanatomyofthegame,ratherthanitsform,controlsthehuntress\'tactics。
Letmemention,beforeIdismissthesubject,asuperbexampleofthismarvellousanatomicaldiscrimination。IoncetookfrombetweenthelegsofaHairyAmmophila,whichhadjustparalysedit,acaterpillarofDicranuravinula。Whatastrangecapturecomparedwiththeordinarycaterpillar!
Bridlinginthickfoldsbeneathitspinkneckerchief,itsfore-partraisedinasphinx-likeattitude,itshinder-partslowlywavingtwolongcaudalthreads,thecuriousanimalisnocaterpillartotheschoolboywhobringsittome,nortothemanwhocomesuponitwhilecuttinghisbundleofosiers;butitisacaterpillartotheAmmophila,whotreatsitaccordingly。Iexplorethequeercreature\'ssegmentswiththepointofaneedle。Allareinsensitive;allthereforehavebeenstung。
CHAPTER12。THEMETHODOFTHESCOLIAE。
AftertheAmmophilae,theparalyserswhomultiplytheirlancet-thruststodestroytheinfluenceofthevariousnerve-centres,exceptingthoseofthehead,itseemedadvisabletointerrogateotherinsectswhichalsoareaccustomedtoanakedprey,vulnerableatallpointssavethehead,butwhichdeliveronlyasinglethrustofthesting。OfthesetwoconditionstheScoliaefulfilledone,withtheirregularquarry,thetenderCetonia-,Oryctes-orAnoxia-larva,accordingtotheScolia\'sspecies。Didtheyfulfilthesecond?Iwasconvincedbeforehandthattheydid。Fromtheanatomyofthevictims,withtheirconcentratednervoussystem,Iforesaw,whencompilingmyhistoryoftheScoliae,thatthestingwouldbeunsheathedonceonly;Ievenmentionedtheexactspotintowhichtheweaponwouldbeplunged。
Thesewereassertionsdictatedbytheanatomist\'sscalpel,withouttheslightestdirectproofderivedfromobservedfacts。Manoeuvresexecutedundergroundescapedtheeye,asitseemedtomethattheymustalwaysdo。
HowindeedcouldIhopethatacreaturewhoseartispractisedinthedarknessofaheapofmouldwoulddecidetoworkinbroaddaylight?Ididnotreckonuponitall。Nevertheless,tosalvemyconscience,ItriedbringingtheScoliaintocontactwithherpreyunderthebell-glass。Iwaswell-advisedtodoso,formysuccesswasininverseratiotomyhopes。
NexttothePhilanthus,noneoftheHuntingWaspsdisplayedsuchardourinattackingunderartificialconditions。Alltheinsectsexperimentedupon,somesooner,somelater,rewardedmeformypatience。LetuswatchtheTwo-
bandedScolia(S。bifasciata,VANDERLIND)operatingonherCetoniagrub。
Theincarceratedlarvastrivestoescapeitsterribleneighbour。Lyingonitsback,itfiercelywendsitswayroundandroundtheglasscircus。
PresentlytheScolia\'sattentionawakensandisbetrayedbyacontinuedtappingwiththetipsoftheantennaeuponthetable,whichnowrepresentstheaccustomedsoil。TheWaspattacksthegame,deliveringherassaultuponthemonster\'shinderend。SheclimbsupontheCetonia-grub,obtainingapurchasewiththetipofherabdomen。Thequarrymerelytravelsthemorequicklyonitsback,withoutcoilingitselfintoadefensiveposture。TheScoliareachesthefore-part,withtumblesandotheraccidentswhichvarygreatlywiththeamountoftolerancedisplayedbythelarva,herimprovisedsteed。Withhermandiblesshenipsapointofthethorax,ontheuppersurface;sheplacesherselfathwartthebeast,archesherselfandmakeseveryefforttoreachwiththeendofherabdomentheregionintowhichthestingistobedriven。Thearchisalittletoonarrowtoembracealmostthewholecircumferenceofhercorpulentprey;andsherenewsherattemptsandeffortsforalongtime。Thetipofthebellytrieseveryconceivableexpedient,touchinghere,thereandeverywhere,butasyetstoppingnowhere。Thispersistentsearchinitselfdemonstratestheimportancewhichtheparalyserattachestothepointatwhichherlancetistopenetratetheflesh。
Meanwhile,thelarvacontinuestomovealongonitsback。Suddenlyitcurlsup;withastrokeofitsheadithurlstheenemytoadistance。
Undiscouragedbyallherset-backs,theWasppicksherselfup,brushesherwingsandresumesherattackuponthecolossus,almostalwaysbymountingthelarva\'shinderend。Atlastafterallthesefruitlessattempts,theScoliasucceedsinachievingthecorrectposition。SheisseatedathwarttheCetonia-grub;themandiblesgripapointonthedorsalsurfaceofthethorax;thebody,bentintoabow,passesunderthelarvaandwiththetipofthebellyreachestheregionoftheneck。TheCetonia-grub,placedinseriousperil,writhes,coilsanduncoilsitself,spinningrounduponitsaxis。TheScoliadoesnotinterfere。Holdingthevictimtightlygripped,sheturnswithit,allowsherselftobedraggedupwards,downwards,sidewards,followingitscontortions。HerobstinacyissuchthatIcannowremovethebell-glassandfollowthedetailsofthedramaintheopen。
Briefly,inspiteoftheturmoil,thetipoftheabdomenfeelsthattherightspothasbeenfound。Thenandonlythenthestingisunsheathed。Itplungesin。Thethingisdone。Thelarva,atfirstplumpandactive,suddenlybecomesflaccidandinert。Itisparalysed。Henceforththerearenomovementssaveoftheantennaeandthemouthparts,whichwillforalongtimeyetbearwitnesstoaremnantoflife。Thepointwoundedhasnevervariedintheseriesofcombatsunderglass:itoccupiesthemiddleofthelineofdemarcationbetweentheprothoraxandthemesothorax,ontheventralsurface。NotethattheCerceres,operatingonWeevils,whosenervoussystemisascompactastheCetonia-grub\'s,driveintheneedleatthesamespot。Similarityofnervousorganizationoccasionssimilarityofmethod。NotealsothattheScolia\'sstingremainsinthewoundforsometimeandrootsaboutwithmarkedpersistence。Judgingbythemovementsofthetipoftheabdomen,onewouldcertainlysaythattheweaponisexploringandselecting。Freetoshiftinonedirectionortheother,withinnarrowlimits,itspointismostprobablyseekingforthelittlemassofnerve-tissuewhichmustbepricked,oratleastsprinkledwithpoison,toobtainoverwhelmingparalysis。
Iwillnotclosethisreportoftheduelwithoutrelatingafewfurtherfacts,ofminorimportance。TheTwo-bandedScoliaisafiercepersecutoroftheCetonia。Inonesittingthesamemotherstabsthreelarvae,oneaftertheother,infrontofmyeyes。Sherefusesthefourth,perhapsowingtofatigueortoexhaustionofthepoison-bag。Herrefusalisonlytemporary。
Nextday,shebeginsagainandparalysestwogrubs;thedayafterthat,shedoesthesame,butwithazealthatdecreasesfromdaytoday。
TheotherHuntingWaspsthatpursuethechasefarafieldgrip,drag,carrytheirprey,afterdeprivingitofmovement,eachinherownfashionand,ladenwiththeirburden,makeprolongedattemptstoescapefromthebell-
glassandtogaintheburrow。Discouragedbythesefutileendeavours,theyabandonthematlast。TheScoliadoesnotremoveherquarry,whichliesonitsbackforanindefinitetimeontheactualspotofthesacrifice。Whenshehaswithdrawnherdaggerfromthewound,sheleaveshervictimwhereitliesand,withouttakingfurthernoticeofit,beginstoflutteragainstthesideoftheglass。Theparalysedcarcaseisnottransportedelsewhere,intoaspecialcellar;therewherethestrugglehasoccurreditreceives,uponitsextendedabdomen,theeggwhencetheconsumerofthesucculenttit-bitwillemerge,thussavingtheexpenseofsettinguphouse。Itgoeswithoutsayingthatunderthebell-glassthelayingdoesnottakeplace:
themotheristoocautioustoabandonhereggtotheperilsoftheopenair。
Whythen,recognizingtheabsenceofherundergroundburrow,doestheScoliauselesslypursuetheCetoniawiththefranticardourofthePhilanthusflingingherselfupontheBee?TheactionofthePhilanthusisexplainedbyherpassionforhoney;hencethemurderscommittedinexcessoftheneedsofherfamily。TheScolialeavesusperplexed:shetakesnothingfromtheCetonia-grub,whichisleftwithoutanegg;shestabs,thoughwellawareoftheuselessnessofheraction:theheapofmouldislackinganditisnothercustomtotransportherprey。Theotherprisoners,oncetheblowisstruck,atleastseektoescapewiththeircapturebetweentheirlegs;theScoliaattemptsnothing。
Afterduereflection,Ilumptogetherinmysuspicionsallthesesurgeonsandaskmyselfwhethertheypossesstheslightestforesight,wheretheeggisconcerned。When,exhaustedbytheirburden,theyrecognizetheimpossibilityofescape,themoreexpertamongthemoughtnottobeginalloveragain;yettheydosobeginafewminuteslater。Thesewonderfulanatomistsknowabsolutelynothingaboutanything,theydonotevenknowwhattheirvictimsaregoodfor。Admirableartistsinkillingandparalysis,theykillorparalyseateveryfavourableopportunity,nomatterwhatthefinalresultasregardstheegg。Theirtalent,whichleavesoursciencespeechless,hasnotashadowofconsciousnessofthetaskaccomplished。
Aseconddetailstrikesme:thedesperatepersistenceoftheScolia。Ihaveseenthestrugglecontinueformorethanaquarterofanhour,withfrequentalternationsofgoodluckandbad,beforetheWaspachievedtherequiredpositionandreachedwiththeendofherabdomenthespotwherethestingshouldpenetrate。Duringtheseassaults,whichwereresumedassoonastheywererepulsed,theaggressorrepeatedlyappliedthetipofherbellytothelarva,butwithoutunsheathing,asIcouldseebytheabsenceofthestartwhichthelarvagiveswhenitfeelsthepainofthesting。TheScoliathereforedoesnotpricktheCetoniaanywhereuntiltheweaponcoverstherequisitespot。Ifnowoundsareinflictedelsewhere,thisisnotinanywayduetothestructureofthelarva,whichissoftandvulnerableallover,exceptinthehead。Thepointsoughtbythestingisnomoreunprotectedthananyotherpartoftheskin。
Inthescuffle,theScolia,curvedintoabow,issometimesseizedbythevice-likegripoftheCetonia-grub,whichisviolentlycoilinganduncoiling。Heedlessofthepowerfulgrip,theWaspdoesnotletgoforamoment,eitherwithhermandiblesorwiththetipofherabdomen。Atsuchtimesthetwocreatures,lockedinamutualembrace,turnoverandoverinamadwhirl,eachofthemnowontop,nowunderneath。Whenitcontrivestoriditselfofitsenemy,thelarvauncoilsagain,stretchesitselfoutandproceedstomakeoffuponitsbackwithallpossiblespeed。Itsdefensiverusesareexhausted。Formerly,beforeIhadseenthingsformyself,takingprobabilityasmyguideIwillinglygrantedtothelarvathetrickoftheHedgehog,whorollshimselfintoaballandsetstheDogatdefiance。
Coileduponitself,withanenergywhichmyfingershavesomedifficultyinovercoming,thelarva,Ithought,woulddefytheScolia,powerlesstounrollitanddisdaininganypointbuttheoneselected。Ihopedandbelievedthatitpossessedthismeansofdefence,ameansbothefficaciousandextremelysimple。Ihadpresumedtoomuchuponitsingenuity。InsteadofimitatingtheHedgehogandremainingcontracted,itflees,bellyinair;
itfoolishlyadoptstheveryposturewhichallowstheScoliatomounttotheassaultandtoreachthespotforthefatalstroke。ThesillybeastremindsmeofthegiddyBeewhocomesandflingsherselfintotheclutchesofthePhilanthus。Yetanotherwhohaslearntnolessonfromthestruggleforlife。
Letusproceedtofurtherexamples。IhavejustcapturedanInterruptedScolia(Colpainterrupta,LATR。),exploringthesand,doubtlessinsearchofgame。Itisamatterofmakingtheearliestpossibleuseofher,beforeherspiritischilledbythetediumofcaptivity。Iknowherprey,thelarvaofAnoxiaaustralis(TheAnoxiaareagenusofBeetlesakintotheCockchafers——Translator\'sNote。);Iknow,frommypastexcavations,thepointsfavouredbythegrub:themoundsofsandheapedupbythewindatthefootoftherosemariesontheneighbouringhill-sides。Itwillbeahardjobtofindit,fornothingisrarerthanthecommonifonewantsitthenandthere。Iappealforassistancetomyfather,anoldmanofninety,stillstraightasacapitalI。Underasunhotenoughtobroilanegg,wesetoff,shoulderinganavvy\'sshovelandathree-prongedluchet。(ThelocalpitchforkofsouthernFrance——Translator\'sNote。)Employingourfeebleenergiesinturns,wedigatrenchinthesandwhereIhopetofindtheAnoxia。Myhopesarenotdisappointed。Afterhavingbythesweatofourbrow——neverwastheexpressionmorejustified——removedandsiftedtwocubicyardsatleastofsandysoilwithourfingers,wefindourselvesinpossessionoftwolarvae。IfIhadnotwantedany,Ishouldhaveturnedthemupbythehandful。Butmypoorandcostlyharvestissufficientforthemoment。To-morrowIwillsendmorevigorousarmstocontinuetheworkofexcavation。